A peer-reviewed open-access journal Zookeys 708: 25—138 (2017) doi: 10.3897/zookeys.708. | 3897 #7,00Ke y S http:/ / ZOO keys -pen soft. net Launched to accelerate biodiversity research Checklist of the freshwater fishes of Colombia: a Darwin Core alternative to the updating problem Carlos DoNascimiento', Edgar Esteban Herrera-Collazos’, Guido A. Herrera-R.*, Armando Ortega-Lara’, Francisco A. Villa-Navarro’, José Saulo Usma Oviedo’, Javier A. Maldonado-Ocampo* | Instituto de Investigacién de Recursos Biolégicos Alexander von Humboldt, Villa de Leyva, Colombia 2 Laboratorio de Ictiologta, Unidad de Ecologta y Sistemdtica (UNESIS), Departamento de Biologia, Facultad de Ciencias, Pontificia Universidad Javeriana, Bogotd, DC, Colombia 3 Grupo de Investigacién en Peces Neotropicales, Fundacién para la Investigacion y el Desarrollo Sostenible (FUNINDES), Cali, Colombia 4 Grupo de Investigacién en Zoologia, Facultad de Ciencias, Universidad del Tolima, Ibagué, Colombia 5 Programa Ecosistemas de Agua Dulce de WWF Colombia, Cali, Colombia Corresponding authors: Carlos DoNascimiento (cdonascimiento@humboldt.org.co); Javier A. Maldonado-Ocampo (maldonadoj@javeriana.edu.co) Academic editor: NV. Bogutskaya | Received 31 May 2017 | Accepted 8 September 2017 | Published 16 October 2017 Attp://zoobank.org/FBOI BD 9E-6656-4FA8-AFB0-4EC546CFF953 Citation: DoNascimiento C, Herrera-Collazos EE, Herrera-R GA, Ortega-Lara A, Villa-Navarro FA, Usma-Oviedo JS, Maldonado-Ocampo JA (2017) Checklist of the freshwater fishes of Colombia: a Darwin Core alternative to the updating problem. ZooKeys 708: 25-138. https://doi.org/10.3897/zookeys.708.13897 Abstract The present work is part of a process to create a Catalogue of the Freshwater Fishes of Colombia and consisted in the depuration and updating of the taxonomic and geographic components of the checklist of the freshwater fishes of Colombia. An exhaustive revision of the 1435 species recorded in 2008 was necessary to: 1. Add new species described since 2009 and species originally described from Colombia but inadvertently omitted in 2008; 2. Add new records of already described species; 3. Delete species whose presence in Colombia was not supported by voucher specimens in ichthyological collections; and 4, Revise the geographic distribution of the species listed in 2008. This process resulted in the following numbers: 1. Total number of freshwater fish species in Colombia: 1494; 2. Number of species recorded by hydrographic region - Amazon: 706, Orinoco: 663, Caribbean: 223, Magdalena-Cauca: 220, Pacific: 130; and 3. Number of endemic species: 374 (76% from the trans-Andean region). Updating the current checklist is a fundamental requirement to ensure its incorporation in the decision-making process with regard to the conservation of Colombian aquatic species and ecosystems, which are facing transformation Copyright Carlos DoNascimiento et al. This is an open access article distributed under the terms of the Creative Commons Attribution License (CC BY 4.0), which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited. 26 Carlos DoNascimiento et al. / ZooKeys 708: 25-138 (2017) processes as a result of activities such as mining, construction of hydroelectric plants, expansion of the agricultural frontier and subsequent deforestation, industrial and domestic pollution, development of waterways, introduction of exotic species, and climate change. Resumen El presente trabajo es parte de un proceso para crear un Catalogo de Peces de Agua Dulce de Colombia y consistié en la depuracién y actualizacién de los componentes taxondémico y geografico del listado de peces de agua dulce de Colombia. Una revisién exhaustiva de las 1435 especies registradas en 2008 fue necesaria para: 1. Adicionar las especies nuevas descritas desde 2009 y especies descritas originalmente para Colombia pero omitidas inadvertidamente en 2008; 2. Adicionar nuevos registros de especies ya descritas; 3. Eliminar especies cuya presencia en Colombia no estaba soportada por especimenes en colec- ciones ictiolégicas; y 4. Revisar la distribucidn geografica de las especies listadas en 2008. Este proceso resulté en las siguientes cifras: 1. Numero total de especies de peces de agua dulce en Colombia: 1494; 2. Numero de especies registradas por regién hidrografica - Amazonas: 706, Orinoco: 663, Caribe: 223, Magdalena-Cauca: 220, Pacifico: 130; y 3. Numero de especies endémicas 374 (76% distribuidas en la region transandina). La actualizacion del presente listado es un requerimiento fundamental para asegurar su incorporacion en el proceso de toma de decisiones en lo concerniente a la conservacién de las especies y ecosistemas acuaticos colombianos, los cuales estan enfrentando procesos de transformacién, como re- sultado de actividades como la minerfa, construccién de plantas hidroeléctricas, expansidn de la frontera agropecuaria y subsecuente desforestacion, polucién industrial y doméstica, desarrollo de hidrovias, intro- duccién de especies exdticas y cambio climatico. Keywords Aquatic ecosystems, conservation, endemic, richness, South America This paper is in honour of the late Richard Peter Vari, who was a co-author of the checklist of the freshwater fishes of Colombia in 2008, a paper that highlighted the importance of the Colombian ichthyofauna in the Neotropical and global contexts, and addressed important issues in terms of the conservation challenges and needs to improve the sampling effort of the continental aquatic systems in Colombia. After almost a dec- ade since that checklist was published, the challenges and tasks pointed out by Richard and co-authors have been attained in several ways in the present paper. He will always be remembered not just for his enormous contribution to the Colombian and Neotropi- cal ichthyology, but most especially for his kindness and willing collaboration. Introduction Since Maldonado-Ocampo et al. (2008), important advances have been made to update the current number of fish species found in Colombian freshwater ecosystems. The first is new fieldwork, which has led to significant range expansions in known distribution of species and new species discoveries; the second is new revisions of existing ichthyo- logical collections. Several checklists have been published at local or regional scales (e.g. Lasso et al. 2009, Villa-Navarro et al. 2011, Ortega-Lara et al. 2012); and an important Checklist of the freshwater fishes of Colombia: a Darwin Core alternative... a7 amount of information has also been published in books that include aspects on ecology, fisheries, threats, and migratory status (e.g. Usma et al. 2009, 2013, Lasso et al. 201 La, Maldonado-Ocampo et al. 2012, Mojica et al. 2012, Zapata and Usma 2013). During this time frame (nine years), fieldwork efforts have been conducted mainly in three of the five hydrographic Colombian regions as delimited by IDEAM (2015): Amazon (Putumayo, Caqueta, Guainia, and Vaupés rivers), Orinoco (piedmont rivers of the Meta and Guaviare basins, and rivers draining directly to the Orinoco as the Bita and Inirida), and Magdalena-Cauca (especially the upper and middle sectors). Some collections in other important rivers as the Catatumbo, Sint, and Atrato (Caribbean system), San Juan and Dagua (Pacific system), have also been conducted. All fishes collected in these rivers are deposited in the following ichthyological collections of Co- lombia (acronyms follow Sabaj, 2016): CAR, CIACOL, CIUA, CP-UCO, CZUT-IC, IAvH-P, ICN-MHN, IMCN, and MPUJ. Complementing these fieldwork and taxonomic efforts that improve the informa- tion quality of the current annotated checklist, an important project “Catalogue of the Freshwater Fishes of Colombia” is in progress. From mid-2014, all previously men- tioned ichthyological collections began a data depuration process. First, and the most important task, all the catalogue data bases associated with these collections are being translated to the Darwin Core standard, allowing the data to be adequately formatted to make them freely accessible through facilitators such as GBIF and SiB Colombia (Colombia Biodiversity Information System: www.sibcolombia.net). At the moment, all available records to mid-2017 in CAR, CIACOL, CICH, CIUA, CP-UCO, CZUT-IC, IAvH-P, ICN-MHN, IMCN, and MPUJ from the hydrographic regions of Orinoco (29000), Magdalena-Cauca (22000), Amazon (14000), Caribbean (5000), and Pacific (1000), have been updated in the following data fields: Occurrence, Organism, Event, Location, Identificacion, and Taxon. The project hopes to conclude this data depuration process for all available records in these collections (around 75000 in total) by December of 2017. The updated annotated checklist presented here and the data available in the collections is the basis for the “Catalogue of the Freshwater Fishes of Colombia” that will be hosted at the Catalogue of Species of the Colombia Biodiver- sity Information System (http://www.biodiversidad.co/#/), and linked to the website of the Colombian Ichthyological Association (ACICTIOS, http://acictios.org). For the first time in Colombia, freshwater ecosystems are the focus of public atten- tion and different strategies are being discussed by the academy and decision-makers to develop adequate conservation and sustainable use strategies for these ecosystems and their biota (Vilardi et al. 2014, Tognelli et al. 2016). The updated information presented here plus the continuous work in the collections to update and edit the data bases will be of extreme importance in terms of the quality and quantity of available information to facilitate evidence-based decision making that can help counter the numerous threats facing aquatic ecosystems (hydropower development, mining, pol- lution, deforestation, exotic species invasion, and climate change scenarios) that are affecting this enormous fish diversity. 28 Carlos DoNascimiento et al. / ZooKeys 708: 25-138 (2017) Methods The starting point for the updating process and corrections to the checklist of the freshwater fishes of Colombia was the checklist of Maldonado-Ocampo et al. (2008). This revision incorporated four main aspects: 1. Addition of newly described species since 2009 and species originally described for Colombia but inadvertently omitted in Maldonado-Ocampo et al. (2008), 2. Addition of new records of already described species, 3. Deletion of nominal species whose presence in Colombia was not supported by voucher specimens in collections, and 4. Revision of the geographic distribution of those species previously listed in Maldonado-Ocampo et al. (2008). The main sources used for this revision consisted in research articles describing new species, taxonomic and systematic revisions of genera and families, direct examination of specimens cata- logued at CZUT-IC, IAvH-P, IMCN, and MPUJ, and to a lesser extent, regional checklists explicitly supported by catalogued lots of specimens for each species entry, and lists of examined material of taxonomic and systematic works dealing with Neo- tropical groups. The complete list of species was assembled using the Darwin Core standard (DwC) and is available both as Suppl. material 1 and through the Integrated Pub- lishing Tool of the Colombian node of GBIF, the SiB Colombia (http://www.sibco- lombia.net/), as version 2.5 at http://doi.org/10.15472/numrso. Future updates of the list will be published in the latter repository. Changes incorporated to each new version of the list will be summarized in the respective metadata section of the elec- tronic resource. A curatorial team presented during the celebration of the biannual meeting of the “Asociacién Colombiana de Ictidlogos” (ACICTIOS) will be in charge of the updating and revision process of the list and of its periodic publication (an- nually or semestrally, depending on the amount of changes accumulated every six months). Below we list fields for which we consider a further detailed explanation of the criteria adopted is needed in order to provide an unequivocal framework for the definitions used for each field. taxonID. This field is defined as a unique and stable-through-time alphanumeric identifier (taxonomic identifier) provided by Life Science Identifier (LSID). namePublishedIn. ‘This field links each species name to its respective source (i.e., a bibliographic reference) and was restricted to include only those species originally described from Colombia, i.e., only the bibliographic reference of the original descrip- tion of species that listed Colombian specimens (in some cases from border regions as well) as types, as well as descriptions that explicitly listed Colombian specimens as non-types. references. [his field is reserved for those species not described from Colombia, but whose presence in the country is supported by a bibliographic reference. In this case, we made a special effort to locate the first chronological reference that explicitly mentioned Colombian specimens collected and deposited in national or international collections or that provided images that allows unambiguous recognition of a determined species, Checklist of the freshwater fishes of Colombia: a Darwin Core alternative... 29 for each main hydrographic system. In some cases, we complemented these primary bibliographic references with more recent references, where additional catalogued lots are provided. Therefore, this approach is somewhat different from the practice imple- mented in Maldonado-Ocampo et al. (2008), in order to avoid the inclusion of refer- ences that are merely lists solely based on previous published works. Future publications reporting new species or distribution extensions of those species already included in this checklist are encouraged to indicate cataloged lots that support them. This action allows traceability and verification of records whenever needed, and facilitates the implementa- tion of new taxonomical arrangements when they need to be proposed. associatedOccurrences. As in the previous field, here we supported the presence of species not originally described from Colombia. We listed catalogued lots (prefer- ably available in Colombian collections) for species not supported by bibliographic references or to complement references that do not provide lots. Lots are provided for each main hydrographic system. In addition, we also listed here catalogued lots cor- responding to types. locationRemarks. Only includes species that are endemic to the Colombian territory. dynamicProperties. Corresponds to the column CT (endangered and threatened species) used in Maldonado-Ocampo et al. (2008). The information in this field was based in Mojica et al. (2012) and since the scope of the present list is of national scale, threat categories are restricted to Colombia. occurrenceRemarks. Indicates the type of commercial use of each species (i.e., Fisheries, Ornamental). The list of fish species included in Colombian inland fisheries is from Lasso et al. (201 1a), while the list of ornamental species is from Ortega-Lara et al. (2015) and Ortega-Lara (2016). locality. Type localities of species described from Colombia are listed as they are cited in their respective original descriptions or as they appear in the Catalogue of Fishes (http://researcharchive.calacademy.org/research/ichthyology/catalog/fishcatmain.asp). Institutional acronyms follow Sabaj (2016). Species formerly indicated as inde- terminate (sp) in Maldonado-Ocampo et al. (2008) are listed at the genus level, since they represent still undescribed species and account for the presence of genera not represented by already described species in Colombia. Classification of taxa of higher taxonomic ranks (Class to Suborder) and arrangement order of respective taxa fol- lowed the proposal of Betancur-R. et al. (2017). The following proposals of classifica- tion were adopted for the indicated groups: Tagliacollo et al. (2016): Gymnotiformes; Mirande (2009, 2010) and Oliveira et al. (2011): Characiformes, but in incongruent cases, we just adopted the latest published proposal (i-e., Oliveira et al. 2011), not implying in any form a validation over the former works of Mirande; Thomaz et al. (2015): Stevardiinae; Sullivan et al. (2006): Siluriformes; Lujan et al. (2015a): Hy- postominae; Roxo et al. (2014): Hypoptopomatinae and Otothyrinae; Covain et al. (2016): Loricariinae; Birindelli (2014): Doradoidea; Sullivan et al. (2013a): Pimelo- doidea; Lépez-Fernandez et al. (2010): Cichlidae. For remaining groups we followed Nelson et al. (2016). 30 Carlos DoNascimiento et al. / ZooKeys 708: 25-138 (2017) Results New species described from Colombia Since the publication of the most recent checklist (Maldonado-Ocampo et al. 2008), 106 new species of freshwater fishes have been described from Colombia (Suppl. ma- terial 2), at a rate of approximately ten species per year. This places the total number of species described or recorded for Colombia at 1494 freshwater fishes (Table 1), a slightly higher figure than the last recorded number of 1435 species (Maldonado- Ocampo et al. 2008). This slight increment in the total number is especially note- worthy since our careful review of each species name in Maldonado-Ocampo et al. (2008) led to the deletion of 202 nominal species that lacked solid support for keep- ing them in the current version of the checklist (see below). The assessment process of some doubtful species led to the inclusion of most of the newly recorded species presented here (see also below), through the direct examination of lots of specimens that were formerly used to support those doubtful species. These figures together, Table |. Number and percentage of species of freshwater fishes by order in Maldonado-Ocampo et al. (2008) compared to the present checklist. Number of species in Number of species Ondes Maldonado-Ocampo et al. (2008) A! in the present work ve Characiformes 637 44.4 644 43.1 Siluriformes 524 36.5 588 39.3 Cichliformes 115 8.0 94 6.3 Gymnotiformes 74 D2, 78 a2 Cyprinodontiformes 35 2.4 42 238 Clupeiformes 11 0.8 10 0.7 Myliobatiformes 9 0.6 11 OF Polycentridae and Sciaenidae (included in Perciformes in 7 0.5 8 0.5 Maldonado-Ocampo et al. 2008) Pleuronectiformes 7 0.5 5 0.3 Beloniformes 5 0.3 4 0.3 Batrachoidiformes fe) 0.2 3 0.2 Osteoglossiformes 3 0.2 3 0.2 Eleotridae (included in Perciformes in 7 0.1 1 0.1 Maldonado-Ocampo et al. 2008) Synbranchiformes 1 0.1 1 0.1 Tetraodontiformes 1 0.1 1 0.1 Ceratodontiformes (Lepidosireniformes 1 0.1 1 01 in Maldonado-Ocampo et al. 2008) 1435 100 1494 100 Checklist of the freshwater fishes of Colombia: a Darwin Core alternative... 31 still place Colombia as the second most-species rich country in the world in terms of its freshwater ichthyofauna. The number of species by hydrographic system is pre- sented in the Table 2, where the Amazon accounts for the highest number of species. A remarkable total of 374 endemic species is now reported from Colombia; 76% of them are species from trans-Andean basins (286 species), and stand as testimony of the unique geological and climatic events that shaped the biogeographic history and taxonomic diversification of the Colombian ichthyological biota. New records of species for Colombia The high number of species already described from other countries but newly recorded for Colombia is a direct result of recent inventories in previously unsampled regions (e.g. Lasso et al. 2009, Villa-Navarro et al. 2011, Ortega-Lara et al. 2012) and perhaps equally or more important, from the verification of the taxonomic identity of records of doubtful species listed in Maldonado-Ocampo et al. (2008), as well as suspicious identifications (i.e., names of species distributed in river basins distant or isolated from the Colombian territory), found in the databases of the examined collections. ‘Thus, the number of new records for the Colombian continental ichthyofauna reaches 114 species (see full list in Suppl. material 3, where also respective hydrographic zone and supporting lot and/or bibliographic reference are provided). Corrections and additions to the geographic distribution of species listed in Mal- donado-Ocampo et al. (2008) The original distribution of 175 species was modified (Suppl. material 4), with 39 species having wider distributions (i.e., encompassing at least one additional hydrographic system than previously recorded), 128 species with more restricted distributions, and 8 species with wrong distributions. At the end of the checklist are Table 2. Number of species by hydrographic system (HS) in Maldonado-Ocampo et al. (2008) com- pared to the present work. Number of species in Number of species in HS Maldonado: Ocamooutal Number of species in the | Maldonado-Ocampo et al. (2008) ? present work (2008) with distribution changes Amazon 788 706 58 Orinoco 658 663 54 Magdalena-Cauca 213 220 40 Caribbean 186 223 iB Pacific 151 130 30 32 Carlos DoNascimiento et al. / ZooKeys 708: 25-138 (2017) summarized the total number of each type of modification (addition or deletion) by hydrographic system. A special treatment was implemented for the species of the genera Astroblepus and Trichomycterus that were originally described from Colombia. Most species of Astroble- pus have restricted geographic distributions, being limited to portions of single river drainages at elevations above 1000 m (Schaefer et al. 2011). A similar scenario is also found for Trichomycterus species, which show high endemism and restricted geographic distributions to headwaters and streams of river systems (Fernandez and Vari 2009, Ferrer and Malabarba 2013, DoNascimiento et al. 2014b). The taxonomic knowledge of the Colombian species of these genera and their respective geographic distributions is precarious. This uncertainty has resulted in misidentifications of specimens that ul- timately result in misplacing species in different disjunct hydrographic zones, without any consideration of the ecological constraints for the geographic distribution of a cer- tain taxonomic group or its biogeographic history. Based on these considerations, we circumscribed the geographic distribution of these species to the hydrographic zone that encompasses the river drainage from which the species were originally described. Species deleted from the Colombian continental ichthyofauna A total of 202 nominal species originally listed in Maldonado-Ocampo et al. (2008) was removed from the updated checklist of the freshwater fishes of Colombia (numeri- cal distribution of deleted species by orders and families is presented in the checklist). These species were found to be absent from Colombian freshwaters or were recently synonymized with other species. Each individual case of species removal is commented (Suppl. material 5) following the format of Lasso et al. (2004). Checklist of the native freshwater fishes of Colombia Abbreviations: DC — described from Colombia. EN — endemic. ET — endangered and threatened species. Amz — Amazon River Basin. Ori — Orinoco River Basin. Mag-Cau — Magdalena-Cauca River Basin. Pac — Pacific slope rivers. Car — Caribbean slope riv- ers. Asterisk (*) denotes species newly recorded for Colombia after Maldonado-Ocam- po et al. (2008). Two asterisks (**) indicate species inadvertently omitted in Maldona- do-Ocampo et al. (2008). Plus (+) and minus (-) signs in the columns corresponding to individual hydrographic systems, indicate respectively, added or deleted presence in each hydrographic system, of species listed in Maldonado-Ocampo et al. (2008) with geographic distribution modified in the present checklist. X indicates unchanged pres- ence in hydrographic systems already accounted in the above bibliographic reference. Checklist of the freshwater fishes of Colombia: a Darwin Core alternative... Table ASD. Number and 33 Taxa DC EN| ET | Amz Mag- Ori Cn Pac| Car References/Collections Myliobatiformes 11 spp (3 added and 2 deleted from 2008) Potamotrygonidae *Heliotrygon gomesi Carvalho & Lovejoy, 2011 ~ Lasso et al. (2013a) Paratrygon aiereba (Miller & Henle, 1841) Plesiotrygon iwamae Rosa, Castello & Thorson, 1987 *Plesiotrygon nana Carvalho & Ragno, 2011 ~ ~ Maldonado-Ocampo et al. (2006a) | Lasso et al. (2013a) Lasso et al. (2010) Lasso et al. (2013a) Potamotrygon constellata (Vaillant, 1880) Potamotrygon magdalenae (Duméril, 1865) Potamotrygon motoro (Miiller & Henle, 1841) ~ 2 Eee va ~ ~ ~ ~ Bogota-Gregory and Maldonado-Ocam- po (2005) Duméril (1865) | Maldonado-Ocampo et al. (2006b) | MNHN A-2368 Mojica et al. (2005) | Maldonado-Ocam- po et al. (2006a) Potamotrygon orbignyi (Castelnau, 1855) ~ Lasso et al. (2013) Potamotrygon schroederi Fernandez-Yépez, 1958 *Potamotrygon scobina Gat- man, 1913 Potamotrygon yepezi Castex & Castello, 1970 x Lasso et al. (2013) Lasso et al. (2013) | Acosta-Santos et al. (2016) Ortega-Lara et al. (2012) Osteoglossiformes 3 spp Osteoglossidae Arapaima gigas (Schinz 1822) Osteoglossum bicirrhosum (Cuvier, 1829) x ~ ~ Mojica et al. (2005) Mojica et al. (2005) | Maldonado-Ocam- po et al. (2006a) Osteoglossum ferreirai Kanaz- awa, 1966 Cala (1977) Clupeiformes 10 spp (1 deleted from 2008) Pristigasteridae 4 spp llisha amazonica (Miranda Ribeiro, 1920) Pellona castelnaeana Valenci- ennes, 1847 Pellona flavipinnis (Valenciennes, 1837) ee ee EE EA ~ a is | ~ ~ Mojica et al. (2005) Correa (2003) | Maldonado-Ocampo sy, (2001) —— Mojica et al. (2005) | Lasso et al. (2005) | ICN-MHN 733, 4144, 11309 34 Carlos DoNascimiento et al. / ZooKeys 708: 25-138 (2017) Taxa DC |EN| ET | Amz| Ori ae Pac| Car| References/Collections Pristigaster cayana Cuvier, il Moi af 1829 ojica et al. (2005) Engraulidae 6 spp (1 deleted from 2008) Amazonsprattus scintilla x Maldonado-Ocampo and Bogota- Roberts, 1984 Gregory (2007) Anchoviella guianensis (Ei- Wee Galvis et al. (2007a) genmann, 1912) ee laa Nas X | X Kullander and Ferraris (2003) peer cede é Majic taboo Lycengraulis batesii (Giinther, ae] al Mojica et al. (2005) | Kullander and 1868) Ferraris (2003) Prerengraulis atherinoides . ee us, 1766) x Galvis et al. (2007a) Characiformes 644 spp (92 added and 93 deleted from 2008) Crenuchidae 28 spp (2 added and 2 deleted from 2008) Crenuchus spilurus Giinther, x |x Arbeléez et al. (2004) | Galvis et al. 1863 (2007a) eee | TL Ex xE | | triacs aoe i pea alg x Weitzman and Kanazawa (1976) 1976 ; FMNH 80401 | USNM 210692, 214364 nae is ea oe, se Mojica et al. (2005) Characidium boavistae Stein- Urbano-Bonilla et al. (2009); Ortega-Lara dachner, 1915 SER si et al. (2012) | Villa-Navarro et al. (2006) Eigenmann (1912) | FMNH 56057 [ex Characidium caucanum xlxlx X CM 4847], 56058-60, 69546 | CAS Eigenmann, 1912 41275-77 [ex IU 12701-03] | USNM 79183 Characidium chupa Schultz, Urbano-Bonilla et al. (2009); Ortega-Lara 1944 et al. (2012) are Mojicaecal. (2005) aia MEA Gi Lae Lasso et al. (2009) | LAvH-P 2753, 13756, phorn, Montafia & Buckup, 13790 2006 Characidium pellucidum Mojica et al. (2005) | Maldonado-Ocam- Eigenmann, 1909 po et al. (2006a) Characidium phoxocephalum x |xlx Eigenmann (1912) | FMNH 56061 [ex Eigenmann, 1912 CM 4851] | CAS 60254 [ex IU 12704] Characidium pteroides Eigen- Maldonado-Ocampo et al. (2008) | mann, 1909 Maldonado-Ocampo et al. (2006a) Checklist of the freshwater fishes of Colombia: a Darwin Core alternative... 35 Taxa DC |EN| ET | Amz| Ori ad Pac| Car| References/Collections Characidium roesseli Géry, x Mojica et al. (2005) 1965 Characidium sanctjohanni x |x Dahl (1960c) | Agudelo-Zamora et al. Dahl, 1960 (2014) Chae ities eter: Urbano-Bonilla et al. (2009). | Bogota- Cope. 1878 Gregory and Maldonado-Ocampo a (2006a) | CAS-SU 50488, 50655 Characidium zebra x |x Arbelaez et al. (2004) | Maldonado- Eigenmann, 1909 Ocampo (2001) Elachocharax geryi Weitzman ; & Kanazawa, 1978 x Weitzman and Kanazawa (1978) a Mal ue x CZUT-IC 4071, 4086, 5106 Weitzman, 1986 ‘ ° Elachocharax pulcher Myers, Mojica et al. (2005) | Weitzman and 1927 Kanazawa (1978) A oat ate Bogota-Gregory and Maldonado- Klausewitzia Géry,1965 Ocampo (2005) Leptocharacidium omospilus RC Meiees IAvH-P 13758 Melanocharacidium dispi- lomma Buckup, 1993 Bacup?) *Melanocharacidium nigrum IAvH-P 8660, 8700, 8765, 8796, 8859, Buckup, 1993 9421 Melanocharacidium pectorale u Buckup, 1993 Buckup (1993) | Mojica et al. (2005) Microcharacidium eleotrioides (Géry, 1960) Maldonado-Ocampo et al. (2008) Microcharacidium gnomus Buckup, 1993 | Buca) Microcharacidium weitzmani Buckup, 1993 Busan iy) Odontocharacidium aphanes (Weitzman & Kanazawa, Buckup (1993) 1977) Erythrinidae 4 spp (1 added and 1 deleted from 2008) Erythrinus erythrinus (Bloch x |x Eigenmann (1922) | Fowler (1945) | & Schneider, 1801) ICN-MHN 440, 3801 Hoplerythrinus unitaeniatus (Agassiz, 1829) Fowler (1945) | Cala (1977) *Hoplias curupira Oyakawa & Mattox 2009 IAVH-P 1349, 2274, 2407, 2743 ; Fowler (1943) | Steindachner (1879a) | aa ROB NEE X_ |Eigenmann (1922) | Maldonado-Ocampo et al. (2013b) | Regan (1914) Parodontidae 9 spp (3 added from 2008) Parodon alfonsoi Londofio- Burbano, Roman-Valencia X Londofo-Burbano et al. (2011) & Taphorn, 2011 36 Carlos DoNascimiento et al. / ZooKeys 708: 25-138 (2017) . | Mag- ; Taxa DC |EN| ET | Amz| Ori Gi Pac| Car| References/Collections Parodon apolinari Myers, Parodon atratoensis Londofio- Burbano, Roman-Valencia Xx. | X X_ |Londofio-Burbano et al. (2011) & Taphorn, 2011 a a Londofio-Burbano et al. (2011) 1887 Parodon caliensis Boulenger, x|xlx Boulenger (1895) 1895 Parodon magdalenensis Londofio-Burbano, Roman- | X | X Londofio-Burbano et al. (2011) Valencia & Taphorn, 2011 Parodon pongoensis (Allen, 1942) Parodon suborbitalis Valenci- ennes, 1850 ~ ~ ~ ~ ~ Londofio-Burbano et al. (2011) Londofio-Burbano et al. (2011) | Eigenmann (1922) Saccodon dariensis (Meek & Hildebrand, 1913) ~ x Pavanelli and Starnes (2015) Cynodontidae 7 spp Cynodon gibbus Spix & Agassiz, 1829 Cynodon septenarius Toledo- Piza, 2000 ~ ~ ~ Mojjica et al. (2005) | Toledo-Piza (2000a) Lasso et al. (2005) | IAVH-P 1043, 1053, 3237 Hydrolycus armatus (Jardine & Schomburgk, 1841) Hydrolycus scomberoides (Cuvier, 1816) ~ ~ Bogota-Gregory and Maldonado- Ocampo (2005) | Maldonado-Ocampo et al. (2006a) Mojica et al. (2005) Hydrolycus tatauaia Toledo- Piza, Menezes & Santos, 1999 Hydrolycus wallacei Toledo- Piza, Menezes & Santos, 1999 ~ ~ TAvH-P 9915 Bogota-Gregory and Maldonado-Ocam- po (2005) | Maldonado-Ocampo et al. (2006a) Rhaphiodon vulpinus Spix & Agassiz, 1829 Serrasalmidae 39 spp (1 added and 4 deleted from 2008) Mojica et al. (2005) | Cala (1977) Catoprion mento (Cuvier, Bogota-Gregory and Maldonado-Ocam- po (2006a) | Lasso et al. (2005) | [AVH-P 1819) Ay | 751, 779,814, 961, 1130, 1173, 1385, 2436, 4911, 5026, 5917, 11272, 12771 | ICN-MHN 5246, 5664 Cilpsictisasradeeasainiine Bogota-Gregory and Maldonado-Ocam- x | x po (2006a) | Cala (1977) | [AVH-P 2031, (Cuvier, 1818) Metynnis argenteus Ahl, 1923 ee ee ee eee ee ee eee eee ~ ~ 5898, 10549, 11601, 11607 Galvis et al. (2007b) | IAVH-P 1399, IAVH-P 11260, [AvVH-P 13864 Checklist of the freshwater fishes of Colombia: a Darwin Core alternative... 37 Taxa Metynnis hypsauchen (Miiller & Troschel, 1844) DC Mag- Cau EN] ET | Amz| Ori Car | References/Collections Bogota-Gregory and Maldonado-Ocam- po (2005) | Maldonado-Ocampo and Bogota-Gregory (2007) Metynnis lippincottianus (Cope, 1870) Bogotd-Gregory and Maldonado-Ocampo (2005) | Maldonado-Ocampo (2001) Metynnis luna Cope, 1878 Myleus rhomboidalis (Cuvier, 1818) Myleus setiger Miller & Troschel, 1844 Myleus torquatus (Kner, 1858) Myloplus asterias (Miller & Troschel, 1844) Correa (2003) | Maldonado-Ocampo (2001) Bogota-Gregory and Maldonado-Ocampo (2005) Bogota-Gregory and Maldonado- Ocampo (2005) | Maldonado-Ocampo et al. (2006a) Mojica (1999) | Maldonado-Ocampo (2001) | ICN-MHN 2557, 3426, 3838 Correa (2003) Myloplus rubripinnis (Miller & Troschel, 1844) Mojica et al. (2005) | Cala (1977) Myloplus schomburgkii (Jardine, 1841) Mylossoma acanthogaster (Valenciennes, 1850) Mylossoma aureum (Spix & Agassiz, 1829) Bogota-Gregory and Maldonado- Ocampo (2005) | Cala (1977) Mojica et al. (2005) Mylossoma duriventre (Cuvier, 1818) Piaractus brachypomus (Cuvier, 1818) Pristobrycon aureus (Spix & Agassiz, 1829) Mojica et al. (2005) | Cala (1977) Mojica et al. (2005) | Cala (1977) Bogotd-Gregory and Maldonado-Ocampo (2006a) | ICN-MHN 5251 Pristobrycon calmoni (Steindachner, 1908) Mojica et al. (2005) Pristobrycon careospinus Fink & Machado-Allison, 1992 Pristobrycon maculipinnis Fink & Machado-Allison, 1992 Pristobrycon striolatus (Steindachner, 1908) Pygocentrus cariba (Hum- boldt & Valenciennes, 1821) Bogota-Gregory and Maldonado- Ocampo (2005) | Maldonado-Ocampo et al. (2006a) Correa (2003) Correa (2003) | Maldonado-Ocampo et al. (2006a) Maldonado-Ocampo (2001) Pygocentrus nattereri Kner, 1858 Mojica et al. (2005) Pygopristis denticulata (Cuvier, 1819) Serrasalmus altuvei Ramirez, 1965 Serrasalmus compressus Jégu, Leao & Santos, 1991 Maldonado-Ocampo et al. (2006a) Correa (2003) | Maldonado-Ocampo (2001) Galvis et al. (2007b) 38 Carlos DoNascimiento et al. / ZooKeys 708: 25—138 (2017) Taxa DC|EN|ET | Amz| Ori ae Pac| Car | References/Collections * Serrasalmus eigenmanni Norman, 1929 Serrasalmus elongatus Kner, CZUT-IC 7334 1858 xX Mojica et al. (2005) Serrasalmus gouldingi (Fink x Bogota-Gregory and Maldonado- & Machado-Allison, 1992) Ocampo (2006a) | IAVH-P 262 Serrasalmus hollandi Eigen- x Mojica et al. (2005) mann, 1915 Serrasalmus humeralis Valen- ciennes, 1850 Bogotd-Gregory and Maldonado-Ocampo (2006a) | ICN-MHN 5255, 6108 Serrasalmus irritans Peters, 1877 Maldonado-Ocampo (2001) Serrasalmus maculatus Kner, 1858 Serrasalmus manueli (Fernan- dez-Yépez & Ramirez, 1967) Serrasalmus medinai Ramirez, 1965 Jégu and Dos Santos (2001) Galvis et al. (2007b) | Maldonado- Ocampo (2001) Mojica et al. (2005) Serrasalmus nalseni Fernandez-Yépez, 1969 Serrasalmus rhombeus (Linnaeus, 1766) Serrasalmus sanchezi Géry, 1964 Correa (2003) IAvH-P 10526-10528, 10539, 10554- 10556, 10559, 11543-11552, 11585 Serrasalmus spilopleura Kner, 1858 Mojica et al. (2005) | Galvis et al. (2007a) Hemiodontidae 14 spp (1 added and 1 deleted from 2008) Anodus elongatus Agassiz, 1829 Mojica et al. (2005) Anodus orinocensis (Stein- dachner, 1887) Hemiodus amazonum (Hum- boldt, 1821) Hemiodus argenteus Pellegrin, 1909 Zapata and Usma (2013) | CUZT-IC 3450, 4062 ICN-MHN 17168 Galvis et al. (2007b) | Galvis et al. (2007a) *Hemiodus atranalis (Fowler, 1940) IAvH-P 2389 Hemiodus gracilis Giinther, 1864 Hemiodus immaculatus Kner, 1858 Hemiodus microlepis Kner, 1858 Maldonado-Ocampo et al. (2006a) Galvis et al. (2007b) | Maldonado- Ocampo et al. (2006a) Arbeléez et al. (2004) | Lasso et al. (2005) | IMCN 606 Hemiodus semitaeniatus Kner, 1858 Hemiodus thayeria Bohlke, 1955 Galvis et al. (2007b) | Cala (1977) Bohlke (1955) | Galvis et al. (2007b) | Ortega-Lara (2016) Checklist of the freshwater fishes of Colombia: a Darwin Core alternative... ay Taxa DC|EN|ET | Amz| Ori ad Pac| Car| References/Collections Hemiodus unimaculatus ; (Bloch, 1794) Galvis et al. (2007b) | Cala (1977) Hemiodus vorderwinkleri : (Géry, 1964) Géry (1964b) Argonectes longiceps (Kner, Maldonado-Ocampo et al. (2006a) | 1858) IAvH-P 5746 Bivibranchia fowleri Galvis et al. (2007b) | Maldonado- (Steindachner, 1908) Ocampo et al. (2006a) Anostomidae 43 spp (4 added and 14 deleted from 2008) . Steindachner (1878) | BMNH oe Asreclae? |x| axa lex x 1895.5.17.155-156 | NMW 69549, ne 69548, 69550 Abramites hypselonotus Lasso et al. (2005) | Mojica et al. (2005) | (Giinther, 1868) IAvH-P 3411 *Anostomoides laticeps : I ; (Eizenmiatn, 1912) XIII Congreso Colombiano de Ictiologia Anostomus anostomus (Lin- Fowler (1943) | Cala (1977) | IAVH-P naeus, 1758) 1014, 2830 Anostomus ternetzi Fernan- dez-Yépez, 1949 les a_i Maldonado-Ocampo et al. (2006a) i ae biders Myers, x Maldonado-Ocampo et al. (2006a) Laemolyta fernandezi Myers, x Galvis et al. (2007a) 1950 Laemolyta garmani (Borodin, x Mojica et al. (2005) 1931) Laemolyta proxima (Garman, iM 1890) Mojica et al. (2005) Laemolyta taeniata (Kner, Mautari and Menezes (2006) | Mojica et 1858) al. (2005) L eee Steindachner (1880) | Garavello and aah i an = 50) Britski (2003) | Maldonado-Ocampo et ae. al. (2005) | Dahl (1955) LeporenusaffinesSetnther, Garavello and Britski (2003) 1864 Leporinus agassizi Steindach- Mojica et al. (2005) ner, 1876 *Leporinus amazonicus : ‘ : Santos & Zuanon, 2008 el | ol XIII Congreso Colombiano de Ictiologia Leporinus aripuanensis 3 Garavello & Santos, 1981 = Mojica etal. (2005) Leporinus bimaculatus bs Ckccitae 188s x Mojica et al. (2005) Garavello (1988) | ANSP 136487, me Wie DOGG Navel SD. las, lige 4 135432-33, 136489, 138803, 140316 | MZUSP 28061 Leporinus brunneus Myers, x |x Galvis et al. (2007b) | Maldonado- 1950 Ocampo et al. (2006a) 40 Carlos DoNascimiento et al. / ZooKeys 708: 25-138 (2017) Taxa DC|EN|ET | Amz| Ori at Pac| Car | References/Collections Leporinus enyae Burns, Chatfield, Birindelli & Burns et al. (2017) Sidlauskas, 2017 Leporinus fasciatus (Bloch, Correa (2003) | Maldonado-Ocampo et 1794) al. (200Ga) Leporinus friderici (Bloch, Mojica et al. (2005) | Maldonado- 1794) Ocampo et al. (2006a) Leporinus granti Eigenmann, x Lasso et al. (2009) | ICN-MHN 948, 1912 1301 Leporinus jamesi Garman, x Garavello et al. (2014) 1929 Leporinus klausewitzi Géry, x Galvis et al. (2007b) 1960 Leporinus moralesi Fowler, x Bogota-Gregory and Maldonado- 1942 Ocampo (2005) Leporinus nattereri Stein- Maldonado-Ocampo et al. (2006a) | dachner, 1876 IAVH-P 8822 nee EPI EROWIEES | allie Fowler (1943) | ANSP 70491, 70492 Leporinus ortomaculatus Garavello, 2000 Bela oee es Up er es pen ay IAvH-P 1822, 12770, 13003, 13011 Leporinus subniger Fowler, Fowler (1943) | LAVH-P 3430-3432 | 1943 ANSP 70493, 70494 Galvis et al. (2007b) | Urbano-Bonilla Leporinus striatus Kner, 1858 X [et al. (2009). | Steindachner (1879a) | Eigenmann (1922) | Regan (1914) Leporinus wolfei Fowler, 1940 nmol 5 [me |] en foi EY Mojica et al. (2005) Eeparenus) appara Eigen, 4 Eigenmann (1922) | CAS 61680 mann, 1922 Megaleporinus muyscorum : ; (Steindachner, 1900) x |X |X x X | Steindachner (1900) | Eigenmann (1922) Megaleporinus trifasciatus . (Steindachner, 1876) = Mojica-ersal-(2005) Pseudanos trimaculatus Kner, x Mojica et al. (2005) 1858 Pseudanos winterbottomi x Neldonadio @icanporerall2006a) Sidlauskas & Santos, 2005 eee mags ears % Rhytiodus argenteofuscus 7. Kner, 1858 Xe Mojica et al. (2005) ne microlepis Kner, | x Mojica et al. (2005) Schizodon corti Schultz, 1944 a ee | Ortega-Lara et al. (2012) Schizodon fasciatus Spix & Agassiz, 1829 x Mojica et al. (2005) Schizodon scotorhabdotus Sidlauskas, Garavello & x Sidlauskas et al. (2007) Jellen, 2007 Checklist of the freshwater fishes of Colombia: a Darwin Core alternative... 4] Taxa DC|EN| ET | Amz| Ori saad Pac| Car| References/Collections Synaptolaemus latofasciatus Maldonado-Ocampo et al. (2006a) | (Steindachner, 1910) CZUT-IC 4092 Chilodontidae 5 spp (1 added and 1 deleted from 2008) Caenotropus labyrinthicus Cala (1977) | Vari et al. (1995) (Kner, 1858) i a eee ie ae ae Bogota-Gregory and Maldonado-Ocam- oe ae a po (2006a) | [AVH-P 281, 4098, 4101 *Caenotropus schizodon Scharcansky & Lucena, CZUT-IC 12280 2007 +p ? Bogota-Gregory and Maldonado-Ocam- ue Lirias Bom po (2006a) | USNM 190300 ,232361, ms 263288, 431461 Chilodus punctatus Miller & R Troschel, 1844 Mojica et al. (2005) | Cala (1977) Curimatidae 47 spp (2 added and 2 deleted from 2008) Curimata aspera (Ginther, : 1868) ¥< Mojica et al. (2005) Curimata cerasina Vari, 1984 = |X CZUT-IC 11494 Curimata cisandina (Allen, a Curimata cyprinoides (Lin- Bogotd-Gregory and Maldonado- naeus, 1766) Ocampo (2006a) Curimata incompta Vati, Mojica et al. (2005) | Maldonado- 1984 Ocampo (2001) | [AvH-P 4370 BI RTE SS all Steindachner (1878) | Vari (1989a) dachner, 1878) Curimata ocellata (Eigen- mann & Eigenmann, 1889) Curimata roseni Vari, 1989 Bogota-Gregory and Maldonado- Ocampo (2006a) Curimata vittata (Kner, 1858) Curimatella alburna (Miller & Troschel, 1844) Curimatella dorsalis (Eigen- mann & Eigenmann, 1889) Vari (1989a) | Maldonado-Ocampo (2001) Arbeldez et al. (2004) Mojica et al. (2005); Maldonado-Ocampo (2001) Curimatella immaculata (Fernandez-Yépez, 1948) Vari (1992b) | Bogota-Gregory and Maldonado-Ocampo (2006a) Curimatella meyeri (Stein- dachner, 1882) Curimatopsis crypticus Vari, 1982 Curimatopsis evelynae Géry, 1964 Mojica et al. (2005) IAvH-P 13266 Géry (1964a) 42 Carlos DoNascimiento et al. / ZooKeys 708: 25-138 (2017) Taxa Curimatopsis macrolepis (Steindachner, 1876) *Curimatopsis microlepis Eigenmann & Eigenmann, 1889 DC EN| ET |Amz|O nears c| Car| References/Collections Vari (1982) Cyphocharax abramoides (Kner, 1858) Cyphocharax aspilos Vati, 1992, Cyphocharax festivus Vari, 1992 Cyphocharax leucostictus (Eigenmann & Eigenmann, 1889) CZUT-IC 4073, 4075 Vari (1992a) X | Ortega-Lara et al. (2012) Ocampo (2006a) | Maldonado-Ocampo Bogota-Gregory and Maldonado- et al. (2006a) Galvis et al. (2007b) Cyphocharax magdalenae (Steindachner, 1878) | Vari (1992a) Cyphocharax multilineatus (Myers, 1927) Cyphocharax nigripinnis Vari, 1992 * Cyphocharax notatus (Steindachner, 1908) Cyphocharax oenas Vari, 1992 Cyphocharax pantostictos Vari & Barriga, 1990 Cyphocharax spiluropsis (Eigenmann & Eigenmann, 1889) Arbeldez et al. (2004) Cyphocharax spilurus (Giinther, 1864) Mojica et al. (2005) | Cala (1977) Potamorhina altamazonica (Cope, 1878) Potamorhina laticeps (Valen- ciennes, 1850) Potamorhina latior (Spix & Agassiz, 1829) Vari (1984) X |Ortega-Lara et al. (2012) Vari (1984) Potamorhina pristigaster (Steindachner, 1876) Psectrogaster amazonica Eigenmann & Eigenmann, 1889 Bogota-Gregory and Maldonado- Ocampo (2006a) Psectrogaster ciliata (Miller & Troschel, 1844) Psectrogaster essequibensis (Giinther, 1864) Psectrogaster rhomboides Eigenmann & Eigenmann, 1889 Mojica et al. (2005) Psectrogaster rutiloides (Kner, 1858) Mojica et al. (2005) Vari (1989b) Mojica et al. (2005) Vari (1989b) Steindachner (1878) | Eigenmann (1922) Galvis et al. (2007b) | Galvis et al. (2007a) CZUT-IC 10874 Maldonado-Ocampo et al. (2006a) IAvH-P 8354-8357 Checklist of the freshwater fishes of Colombia: a Darwin Core alternative... 43 Taxa DC|EN| ET | Amz| Ori ad Pac| Car| References/Collections Pseudocurimata lineopunctata Se) ei | Boul@neeriQOii),| Higenmanns 92) (Boulenger, 1911) ee ania Pseudocurimata patiae x |x x Eigenmann et al. (1914) | FMNH 56554, (Eigenmann, 1914) 56555 | CAS 60622 [ex IU 13055] Steindachnerina argentea (Gill, 1858) r+) | 8S Eigenmann (1922) Eigenmann (1912) | FMNH 56024 [ex Steindachnerina atratoensis x |x x CM 48 14a], 56043, 71308 | CAS 44218 (Eigenmann, 1912) [ex IU 12676] | MCZ 30933 | USNM FILS? Steindachnerina bimaculata ; (Steindachner, 1876) a van te) Steindachnerina dobula ine SL. |_| lieiteas Steindachnerina guentheri (Eigenmann & Eigenmann, DEO ee Arbelaez et al. (2004) | Vari (1991) 1889) Steindachnerina hypostoma a (Boulenger, 1887) x Mojica et al. (2005) Steindachnerina pupula Vari, x Vari (1991) 1991 Prochilodontidae 10 spp Ichthyoelephas longirostris x |x x ‘ Steindachner (1879b) | Castro and Vari (Steindachner, 1879) (2004) Prochilodus magdalenae x |x x x Steindachner (1879a) | Eigenmann Steindachner, 1879 (1922) Prochilodus mariae Biderimant 199) x Eigenmann (1922) Prochilodus nigricans Spix & x Fowler (1943) Agassiz, 1829 Prochilodus reticulatus Valenciennes, 1850 Prochilodus rubrotaeniatus Jardine & Schomburgk, 1841 Semaprochilodus insignis (Jar- dine & Schomburgk, 1841) Semaprochilodus kneri (Pellegrin, 1909) Xx x Ortega-Lara et al. (2012) Bogota-Gregory and Maldonado- Ocampo (2006a) | Urbano-Bonilla et al. (2009). | IAVH-P 5844, 6195, 10879, 10880, 12764 Mojica et al. (2005) | Castro and Vari (2004) Correa (2003) | Maldonado-Ocampo et al. (200Ga) Semaprochilodus laticeps (Steindachner, 1879) x Cala (1977) | Castro and Vari (2004) Semaprochilodus taeniurus (Valenciennes, 1821) Lebiasinidae 30 spp (3 added and 10 deleted from 2008) Correa (2003) Lebiasina astrigata Regan, 1913 ee eee ee ee ee aE! Xx Maldonado-Ocampo et al. (2006b) 44 Carlos DoNascimiento et al. / ZooKeys 708: 25-138 (2017) Taxa Lebiasina chocoensis Ardila DC Ori ~) 6 Car References/Collections Rodriguez, 2008 Lebiasina panamensis Gill, 1877 Copeina guttata (Steindachner, 1876) Xx 4200, 4201 Eigenmann (1922) | Fowler (1944) Steindachner (1876) Rodriguez, 2010 x | X lethal les Ardila Rodriguez (2010) Lebiasina chucuriensis Ardila x |x x Ardila Rodriguez (2001) | CAR 150424, Rodriguez, 2001 150423, 150425 Lebiasina colombia Ardila x |x x Ardila Rodriguez (2008b) | CAR 190, Rodriguez, 2008 191, 192 | ICN-MHN 5314 Lebiasina elongata : (Boule os 37) x Galvis et al. (2007b) Maldonado-Ocampo et al. (2005) | Lebiasina erythrinoides Lasso et al. (2005) | Bogota-Gregory and (Valenciennes, 1850) Maldonado-Ocampo (2005) | Ortega- Lara et al. (2012) | IAVH-P 573 Lebiasina festae Boulenger, Maldonado-Ocampo et al. (2013b) | 1899 Fowler (1939) Lebiasina floridablancaensis Ardila Rodriguez (1994) | CAR 15-04-01 Ardila Rodriguez, 1994 X |X xX | MBUCV-V-26713 [ex CAR 15-04-02 and 03] | MHNUNC 1837 Lebiasina multimaculata x |x x Boulenger (1911) | BMNH Boulenger, 1911 1910.7.11.167-169 Lebiasina narinensis Ardila x |x Ardila Rodriguez (2002) | ICN-MHN Rodriguez, 2002 2340, 2340-1 Vebiaind oregiéardila Tat fl Ardila Rodriguez (2008a) | CAR 157, 265 x |X X | CZUT-IC 2586 | [AvH-P 9875 | IMCN xX xX Copella eigenmanni (Regan, 1912) Regan (1912d) Copella nattereri (Steindach- ner, 1876) Copella vilmae Géry, 1963 Pyrrhulina brevis Steindach- ner, 1876 Pyrrhulina eleanorae Fowler, 1940 ~ ~ Arbeldez et al. (2004) | Galvis et al. (2007a) Mojica et al. (2005) Correa (2003) | Urbano-Bonilla et al. (2009). Maldonado-Ocampo et al. (2008) Pyrrhulina filamentosa Valenciennes, 1847 Pyrrhulina laeta (Cope, 1872) Pyrrhulina lugubris Eigen- mann, 1922 ~ x |X ~ ~ ~< Lasso et al. (2005) | IAVH-P 1589, 3405 Arbeldez et al. (2004) Eigenmann (1922) | CAS 78888 [ex IU 15041a], 78888 [ex IU 15041] Pyrrhulina obermulleri Myers, 1926 Mojica et al. (2005) Pyrrhulina semifasciata Stein- dachner, 1876 *Nannostomus digrammus (Fowler, 1913) Bogoté-Gregory and Maldonado-Ocampo (2006a) | USNM 311003 TAvH-P 10688, 10692, 10694 Checklist of the freshwater fishes of Colombia: a Darwin Core alternative... 45 Taxa Nannostomus eques Stein- dachner, 1876 Nannostomus harrisoni (Eigenmann, 1909) DC EN | ET | Amz Car | References/Collections Mojica et al. (2005) | Maldonado- Ocampo et al. (2006a) Bogota-Gregory and Maldonado- Ocampo (2005) | Maldonado-Ocampo et al. (2006a) Nannostomus marginatus Eigenmann, 1909 Nannostomus marilynae Weitzman & Cobb, 1975 Nannostomus trifasciatus Steindachner, 1876 Weitzman (1966) | Lasso et al. (2005) Weitzman and Cobb (1975) Arbelaez et al. (2004) | Galvis et al. (2007a) Nannostomus unifasciatus Steindachner, 1876 Ctenoluciidae 7 spp Boulengerella cuvieri (Agas- siz, 1829) Arbeldez et al. (2004)Maldonado- Ocampo et al. (2006a) Vari (1995) Boulengerella lateristriga (Boulenger, 1895) Correa (2003) | Maldonado-Ocampo et al. (2006a) Boulengerella lucius (Cuvier, 1816) Boulengerella maculata (Valenciennes, 1850) Maldonado-Ocampo et al. (2006a) Vari (1995) | Cala (1977) Boulengerella xyrekes Vari, 1995 Ctenolucius beani (Fowler, 1907) Ctenolucius hujeta (Valenciennes, 1850) Bogota-Gregory and Maldonado- Ocampo (2006a) | Maldonado-Ocampo et al. (2006a) | IAVH-P 1789, 2044, 2851 Fowler (1907) | Vari (1995) X | Vari (1995) | Bean (1908) Acestrorhynchidae 18 spp (1 added and 1 deleted from 2008) Acestrorhynchus abbreviatus (Cope, 1878) Acestrorhynchus falcatus (Bloch, 1794) Galvis et al. (2007b) Galvis et al. (2007b) | Eigenmann (1922) Acestrorhynchus falcirostris (Cuvier, 1819) Acestrorhynchus grandoculis Menezes & Géry, 1983 Acestrorhynchus heterolepis (Cope, 1878) Mojica et al. (2005) | Maldonado- Ocampo et al. (2006a) Maldonado-Ocampo et al. (2006a) Lasso et al. (2005) | Galvis et al. (2007b) | IAvH-P 834, 918, 2157, 10772 Acestrorhynchus microlepis (Schomburgk, 1841) Mojica et al. (2005) | Toledo-Piza and Menezes (1996) | Cala (1977) Acestrorhynchus minimus Menezes, 1969 Acestrorhynchus nasutus Eigenmann, 1912 Maldonado-Ocampo et al. (2006a) Lasso et al. (2009) | IAVH-P 1124, 9908, 9969;.9910; 9911, 9912,99 13,9914, 13688, 13797, 13875 | ICN-MHN 1280 46 Carlos DoNascimiento et al. / ZooKeys 708: 25-138 (2017) . | Mag- ; Taxa DC | EN| ET | Amz| Ori Gi Pac| Car | References/Collections Gnathocharax steindachneri Fowler, 1913 ak ane Teena ares eer Toledo-Piza (2000b) | LAvH-P 10658 Eigenmann, 1912 * Heterocharax virgulatus Toledo-Piza, 2000 ee Hoplocharax goethei Géry, IAVEL-P 15984 1966 Lonchogenys ilisha Myers, Mojica et al. (2005) | Maldonado- 1927 Ocampo et al. (2006a) Cueriesiains x Steindachner (1878) | Eigenmann (1922) (Steindachner, 1878) hie Sree Se tenias Boanaees x X | Schultz (1943) | USNM 76976, 120170 Schultz, 1943 Gilbertolus maracaiboensis Schultz, 1943 Roestes molossus (Kner, 1858) Roestes ogilviei (Fowler, 1914) Characidae 325 spp (68 added and 51 deleted from 2008) Brachychalcinus copei (Steindachner, 1882) Brachychalcinus nummus Bohlke, 1958 Ortega-Lara et al. (2012) Prada-Pedreros (1997) Galvis et al. (2007b) Mojica et al. (2005) Bogota-Gregory and Maldonado- Ocampo (2005) Brachychalcinus orbicularis (Valenciennes, 1850) Mojica et al. (2005) | ICN-MHN 1309 | 824, 839, 862, 865 Gymnocorymbus bondi (Fowler, 1911) Gymnocorymbus thayeri Eigenmann, 1908 Benine et al. (2015) Fowler (1945) | Galvis et al. (2007a) x be x 2.6 = i i Poptella brevispina Reis, 1989 xX TAvH-P 4382 Poptella compressa (Giinther, x Mojica et al. (2005) | Maldonado- 1864) a Ocampo and Bogotd-Gregory (2007) ew longepannss Popes, x Maldonado-Ocampo et al. (2013a) Stethaprion erythrops (Cope, x Galvis et al. (2007b) 1870) Stichonodon insignis 4 (Steindachner, 1876) * Mojicasecal (2005) soe st dal (ES X | |Dahl (1960c) | ICN-MHN 147, 147a en BS Eigenmann (1912) | FMNH 56022 [ex ae ta ra Be |X X | X |CM 4812], 56023, 69678 | CAS 57600 acest [ex IU 12675] | USNM 79225 se Mase ae Sl ss x Steindachner (1880) | NMW 57983 (Steindachner, 1880) Roeboides araguaito Lucena, Checklist of the freshwater fishes of Colombia: a Darwin Core alternative... 47 Taxa DC|EN| ET | Amz| Ori oad Pac| Car| References/Collections Acestrocephalus boehlkei ; Bogota-Gregory and Maldonado-Ocampo Acestrocephalus sardina x |x (2006a) | Lasso et al. (2005) | [AvH-P (Fowler, 1913) 4092-4093, 4793-4796,5851,12001, 12025, 12026, 12119 Charax condei (Géry & y Eigenmann (1922) | CAS 41300 [ex IU 15027], 69117 [ex IU 15027] | FMNH Charax metae Eigenmann, x 55145 | IAVH-P 3566, 3567, 3569, 1922. 3571, 3573, 3574, 7732 | SU 60707 [ex IU 15028] | UMMZ 160226 | USNM 83631 [ex IU 15027] Charax michaeli Lucena, x Mojica et al. (2005) 1989 Charax niger Lucena, 1989 a ene Mojica et al. (2005) ae notulatus Lucena, | x IAVH-P 7521, 13028, 13073 Charax tectifer (Cope, 1870) | re a Arbeldez et al. (2004) Cynopotamus amazonus ¥ (Giinther, 1868) xX Mojica et al. (2005) Cynopotamus atratoensis x |xlx x Eigenmann and Ogle (1907) | USNM (Eigenmann, 1907) 1664, 306567 [ex USNM 1664] Cynopotamus bipunctatus Pellegrin, 1909 x Urbano-Bonilla et al. (2009). Cynopotamus magdalenae x|xlx x Steindachner (1879a) | NMW 62501-02, (Steindachner, 1879) 62504-05, 77769 Cynopotamus venezuelae (Schultz, 1944) X | Ortega-Lara et al. (2012) Exodon paradoxus Miller & Troschel, 1844 . Cala (1977) Galeocharax gulo (Cope, 1870) sa Tt Mojica et al. (2005) *Phenacogaster maculoblonga de Lucena & Malabarba, XxX CZUT-IC 9680 2010 *Phenacogaster napoatilis de Lucena & Malabarba, 2010 sa iid Phenacogaster pectinata 4 *Phenacogaster prolata de Lucena & Malabarba, 2010 x a pele GAs, weneNeta Weitzman and Vari (1987) | NRM 15048 case ee ||| ex x | MBUCV-V-15342 | MZUSP 36498 | ee NRM 17593 | USNM 278479 Roeboides affinis (Giinther, x |x Lucena (2007) 1868) x 2003 Lucena (2003) 48 Carlos DoNascimiento et al. / ZooKeys 708: 25-138 (2017) Taxa DC|EN|ET | Amz| Ori ae Pac| Car | References/Collections Roeboides dayi (Steindachner, 1878) Roeboides dientonito Schultz, 1944 Roeboides myersii Gill, 1870 Roeboides occidentalis Meek & Hildebrand, 1916 Steindachner (1878) | Maldonado- Ocampo et al. (2013b) Lasso et al. (2005) | Ortega-Lara et al. (2012) | IAVH-P 2935, 2988, 3329-3332, + |4906, 9497, 13083, 13169, 13185, 13198, 13216, 13236, 13249, 13278, 13296, 13376 Mojica et al. (2005) Maldonado-Ocampo et al. (2013b) Bario steindachneri (Eigenmann, 1893) Hemigrammus aguaruna Lima, Correa & Ota, 2016 Hemigrammus analis Durbin, 1909 Galvis et al. (2006) Lima et al. (2016) | Galvis et al. (2006) Arbelaez et al. (2004) | Maldonado- Ocampo (2001) Hemigrammus barrigonae Eigenmann & Henn, 1914 Hemigrammus bellottii (Steindachner, 1882) Hemigrammus bleheri Géry & Mahnert, 1986 Eigenmann and Henn (1914) | AMNH 5320 | CAS 44368 [ex IU 13423], 44369 [ex IU 13424] | IAVH-P 3633 Mojica et al. (2005) | Cala (1977) | IAvH-P 10304, 10307 Géry and Mahnert (1986) Hemigrammus coeruleus Durbin, 1908 *Hemigrammus geisleri Zatske & Géry, 2007 Hemigrammus gracilis (Liitken, 1875) Hemigrammus hyanuary Durbin, 1918 Bogota-Gregory and Maldonado-Ocampo (2006a) | ICN-MHN 5065 CZUT-IC 4400, 5097, 8655, 8851, 8933, 8954, 9099, 9245, 9993, 14642, 16138, 16309, 16837, 16934, MPUJ 306, 1232, 1234, 1237, 1357, 4378, 4392, 10804 Maldonado-Ocampo (2001) | Fowler (1945) | IAvH-P 8735, 8763 Castro (1987a) | Lasso et al. (2005) | ICN-MHN 2787 Hemigrammus levis Durbin, 1908 Correa (2003) | Maldonado-Ocampo et al. (2006a) Hemigrammus luelingi Géry, 1964 Hemigrammus lunatus Mojica et al. (2005) Durbin, 1918 x Mojica et al. (2005) Hemigrammus melanochrous Fowler, 1913 = Cortea (2005) Hemigrammus micropterus Meek, 1907 X Maldonado-Ocampo et al. (2006a) ae se, Bogota-Gregory and Maldonado- ake ate ae ar x [x Ocampo (2006a) | Cala (1977) | LAVH-P ik: 2605 | IICN-MHN 4273 Hemigrammus newboldi x |x Maldonado-Ocampo (2001) | CZUT-IC (Fernandez-Yépez, 1949) 7204, 7250, 14763 Checklist of the freshwater fishes of Colombia: a Darwin Core alternative... 49 Taxa DC |EN| ET | Amz| Ori ae Pac| Car| References/Collections Hemigrammus ocellifer x Mojica (1999) | CLACOL 2327, CZUT- (Steindachner 1882) IC 18072 ir five bn Bogota-Gregory and Maldonado-Ocampo anes aaa i aa x (2006a) | USNM 216881 | ICN-MHN: == 16850-16851 Hemigrammus pulcher . Ladiges, 1938 2S Mojica et al. (2005) Hemigrammus rhodostomus x lx Galvis et al. (2007b) | Cala (1977) Ahl, 1924 Mer ae - Hemigrammus rubrostriatus x |x x Zarske (2015) Zarske, 2015 Hemigrammus schmardae (Steindachner, 1882) Hemigrammus stictus (Durbin, 1909) Hemigrammus unilineatus (Gill, 1858) ~ ~ : Arbelaez et al. (2004) | Maldonado- Ocampo et al. (2006a) Cala (1977) Galvis et al. (2007b) | Cala (1977) Hemigrammus vorderwinkleri Géry, 1963 Correa (2003) | Maldonado-Ocampo et al. (2006a) *Hemigrammus yinyang Lima & Sousa, 2009 Hyphessobrycon acaciae Garcia-Alzate, Roman- CZUT-IC 4323, 4467, 4553, 4857, MPUJ 12525, 12623-12631 Garcfa-Alzate et al. (2010a) | MPUJ 5683, Regan, 1913 Hyphessobrycon copelandi Durbin, 1908 Mojica et al. (2005) Hyphessobrycon diancistrus Weitzman, 1977 *Hyphessobrycon dorsalis Zarske, 2014 Weitzman (1977) | USNM 216607, 216606 | BMNH 1977.1.12.1-2 | MZUSP 13179-80 IAvH-P 1032, 2315 Valencia & Prada-Pedreros, Shee * 393 | IUQ 2795, 2433, 2492, 2793 2010 Bogota-Gregory and Maldonado- Ocampo (2006a) | [AVH-P 8331-8332, Hyphessobrycon agulha x 8334, 8336-8344, 8346-8350, 8728, Fowler, 1913 8794, 8888, 8919, 8941, 8953, 8973, 8989, 9012, 9025, 9046, 9071, 9096, 9121, 9407, 11175-11178 Hyphessobrycon amaronensis Garcia-Alzate, Roman- x |X X Garcfa-Alzate et al. (2010b) | IUQ 2286 Valencia & Taphorn, 2010 Hyphessobrycon bentosi ae |e Bogota-Gregory and Maldonado- Durbin, 1908 Ocampo (2005) Hyphessobrycon chocoensis Garcia-Alzate et al. (2013) | IUQ 3035 | Garcfa-Alzate, Roman- x | Xx xX AUM 55075 | IUQ 2274, 2275, 3036, Valencia & Taphorn, 2013 3044 Hyphessobrycon columbianus x |x x Zarske and Géry (2002) | MTDF 25497, Zatske & Géry, 2002 25498 Epp pesobryeare condones © | $0 | Regan (1913) | BMNH 1913.10.1.19-21 x x 50 Carlos DoNascimiento et al. / ZooKeys 708: 25-138 (2017) . | Mag- ; Taxa DC |EN| ET | Amz| Ori ene Pac| Car| References/Collections *Hyphessobrycon epicharis Weitzman & Palmer, 1997 ti Se ee Hyphessobrycon erythrostigma ; (Fowler, 1943) x Weitzman and Palmer (1997) Hyphessobrycon gracilior x Bogota-Gregory and Maldonado-Ocampo Géry, 1964 (2006a) | ICN-MHN 4277 *Hyphessobrycon } heterorhabdus (Ulrey, 1894) 2s Garcfa-Alzate et al. (2008) Hyphessobrycon klausanni Garcia-Alzate, Urbano- x |X X Garcia-Alzate et al. (2017) Bonilla & Taphorn, 2017 Ebpbespe yonre lores Xx ICN-MHN 16814, 16846-16848 Ladiges, 1938 Hyphessobrycon mavro Garcia-Alzate, Roman- x |x x Garcfa-Alzate et al. (2010a) | IUQ 2791, Valencia & Prada-Pedreros, 1964 | IMCN 3751 2010 Hyphessobrycon melazonatus x Castro and Arboleda (1988) | ICN-MHN Durbin, 1908 16855-16856 Hyphessobrycon metae ; Bigenimann oe tean, 104d x: Eigenmann and Henn (1914) Hyphessobrycon minimus Durbin, 1909 X Maldonado-Ocampo et al. (2008) Hyphessobrycon natagaima Garcia-Alzate, Taphorn, : Romén-Valencia & Villa- Km, || EX XxX Garcia-Alzate et al. (2015b) Navarro, 2015 Hyphessobrycon niger Garcia- F Alzate, Roman-Valencia& | X | X X = SSO ea | MPU Prada-Pedreros, 2010 ; Hyphessobrycon ocasoensis f ; ; Coca Ribate Be Rorabe x |x x oe and Roman-Valencia Valencia, 2008 Hyphessobrycon oritoensis Garcia-Alzate et al. (2008) | IUQ 1574, Garcfa-Alzate, Roman- X |X xX 129, 1575 | MBUCV-V 33737 | MCGN Valencia & Taphorn, 2008 55844 *Hyphessobrycon otrynus Benine & Lopes, 2008 a On aett) Hyphessobrycon peruvianus - Ladiges, 1938 xX Mojica et al. (2005) ah Eigenmann (1913) | CAS 77396 [ex IU ce pass ad x |x| x calle 12850] | FMNH 56290 [ex CM 5091], oe aes 56291, 75150 | USNM 79214 Eigenmann (1913) | Garcia-Alzate et al. (2015b) | CAS 57603 [ex IU 12858], Hyphessobrycon proteus x |x x x 60478-83 [ex IU 12852-57] | CIUA 295, Eigenmann, 1913 296 | FMNH 56293 [ex CM 5094], 56294-99, 56381-82, 69778 | SU 22754 | USNM 79215, 167819. Hyphessobrycon saizi Géry, x |x x Géry (1964a) | USNM 198647 | ZMA 1964 114206 Checklist of the freshwater fishes of Colombia: a Darwin Core alternative... 51 Taxa DC|EN| ET |Amz| Ori ae Pac| Car | References/Collections Hyphessobrycon sebastiani Garcia-Alzate, Roman- ee |e x Garcia-Alzate et al. (2010c) Valencia & Taphorn, 2010 Ree EDS X | Ortega-Lara et al. (2012) Hyphessobrycon sweglesi x |x _|x Géry (1961) | USNM 196090, IAVH-P (Géry, 1961) 9517, 11276 Hyphessobrycon taguae Cottiacil ; ; ‘ arcia-Alzate et al. (2010b) | Garcfa- Garcia-Alzate, Roman- ee | > an 4 Alzate et al. (2010a) | TUQ 2288 Valencia & Taphorn, 2010 Hyphessobrycon tukunai Géry, x Bogota-Gregory and Maldonado-Ocampo 1965 (2006a) | ICN-MHN 4278 Moenkhausia browni Je Galvis et al. (2007b) | Maldonado- Eigenmann, 1909 Ocampo (2001) ee fogs Xx Maldonado-Ocampo et al. (2006a) Meeabnacyaaien Maldonado-Ocampo (2001) | IAVH-P (Giinther, 1864) el | EX 8652, 8690, 8725, 8766, 8797, 8830, 8860, 8914 Necetieag ales Sfx fx} || feos 2003) | cata 1977 eae x Mojica etal. (2005) Bogota-Gregory and Maldonado- Rid ea a 2) x | Ban (20062) | Maldonado-Ocampo ; et al. (2004) | ICN-MHN 11028, 11130 Moenkhausia cotinho x |x Galvis et al. (2007b) | Maldonado- Eigenmann, 1908 Ocampo and Bogotd-Gregory (2007) Moenkhausia dichroura x |x Mojica et al. (2005) | Maldonado- (Kner, 1858) Ocampo et al. (2006a) ie ee Géry (1964a) | ANSP 112249, 139712 | Géry, 1964 XxX | X& xX ICN-MHN 17133 | MNHN 1980-1445 | USNM 198640 teri casi erltadicgittiolt Mojica (1999) | Maldonado-Ocampo et (Miiller & Troschel, 1845) X | X al. (2006a) | ICN-MHN 3838, 6035, 6091 *Moenkhausia hysterosticta Lucinda, Malabarba & X CZUT-IC 9821, MPUJ 4387, 4468 Benine, 2007 Moenkhausia intermedia x |x Galvis et al. (2007b) | Maldonado- Eigenmann, 1908 Ocampo et al. (2006a) ea ese an x Maldonado-Ocampo et al. (2006a) *Moenkhausia justae eee ee x IAVH-P 11014-11015 Moenkhausia lepidura (Kner, Arbelaez et al. (2004) | Maldonado- 1858) Ocampo et al. (2006a) Moenkhausia megalops Arbeléez et al. (2004) | Maldonado- (Eigenmann, 1907) Ocampo et al. (2006a) 52 Carlos DoNascimiento et al. / ZooKeys 708: 25-138 (2017) Taxa DC |EN| ET | Amz) O ae "| Pac| Car] References/Collections Rae ane px f=] || faster ea. 2008 ial Eigenmann (1922) | CAS 55610 [ex IU Moenkhausia metae x 15026a], 55609 [ex IU 15026], 55608 Eigenmann, 1922 [ex IU 13951] | FMNH 55214 | IAvH-P 9177 *Moenkhausia mikia CZUT-IC 3498, 3555, 3584, 4910, Marinho & Langeani, 2010 4918, 5104, 8069 Rnguena psa aligegnis Fowler (1943) | Eigenmann (1922) (Giinther, 1864) es eee x Fowler (1943) | ANSP 70496 Moenkhausia robertsi Géry, Bogota-Gregory and Maldonado-Ocampo 1964 (2006a) | ICN-MHN 5511, 6036 Moenkhausia simulata (Eigenmann, 1924) Correa (2003) Paracheirodon axelrodi (Schultz, 1956) Cala (1977) Paracheirodon innesi (Myers, 1936) Paracheirodon simulans (Géry, 1963) Parapristella georgiae Géry, 1964 Petitella georgiae Géry & Boutiére, 1964 Mojica et al. (2005) Weitzman and Fink (1983) Géry (1964a) | Lasso et al. 2004, Maldonado-Ocampo 2004, Maldonado- Ocampo et al. 2006b, Maldonado- Ocampo et al. 2008, Lasso et al. 2009 Galvis et al. (2007b) Thayeria obliqua Eigenmann, 1908 Maldonado-Ocampo (2001) | ICN- MHN 6025, 17140 Tetragonopterus argenteus Cuvier, 1816 Tetragonopterus chalceus Spix & Agassiz, 1829 Aphyocharax alburnus (Giinther, 1869) Mojica et al. (2005) | Fowler (1945) Fowler (1943) | Cala (1977) Fowler (1943) | Maldonado-Ocampo et al. (2006a) “Aphyocharax colifax Taphorn & Thomerson, 1991 Aphyocharax erythrurus Eigenmann, 1912 Aphyocharax pusillus Giinther, 1868 CAITAIG A271. Galvis et al. (2007b) | ICON-MHN 5387 Galvis et al. (2007b) Paragoniates alburnus Steindachner, 1876 Mojica et al. (2005) | Galvis et al. (2007a) Phenagoniates macrolepis (Meek & Hildebrand, 1913) Prionobrama filigera (Cope, 1870) Xenagoniates bondi Myers, 1942 Eigenmann and Henn (1914) | Eigenmann (1922) Mojica et al. (2005) Gala(1977) Checklist of the freshwater fishes of Colombia: a Darwin Core alternative... a5 Taxa Atopomesus pachyodus Myers, 1927 Axelrodia riesei Géry, 1966 DC Car | References/Collections Myers (1927) | CAS 41736 [ex IU 17673] Géry (1966a) | USNM 207923, 207924 | Axelrodia stigmatias (Fowler, 1913) Microschemobrycon callops Bohlke, 1953 Microschemobrycon casiquiare Bohlke, 1953 xX ANSP 112512, 139715 | MNHN 1982- 0538 | ZMA 113865 x Arbeldez et al. (2004) Maldonado-Ocampo et al. (2006a) Galvis et al. (2007b) | Maldonado- Ocampo et al. (2006a) Microschemobrycon geisleri Géry, 1973 *Microschemobrycon melanotus (Eigenmann, 1912) *Oxybrycon parvulus Géry, 1964 Arbeldez et al. (2004) MPUJ 236, 247, 630, 1436, 4497 CZUTAIG 47910517953 *Parecbasis cyclolepis Eigenmann, 1914 CIACOL 272 * Tyttobrycon xeruini Géry, 1973 TAvH-P-14408, 15186, 15208, 15216, 15233, 15244, 15256, 15358, 15391, 15427, 15433, 15469, 15482, 15517, 15556, 15573, 15597, MPUJ 11025 Grundulus bogotensis Humboldt and Valenciennes (1821) | (Humbolde, 1821) Sill ZMB 33306, 3505, 31499 ee ee sel ak Romén-Valencia etal. (2001)| 1UQ 453, ; ‘ 454, 455 | STRI 1369 2003 Weitzman and Fink (1971) | USNM Nematobrycon lacortei x |x 205594, 205595-96 | BMNH Weitzman & Fink, 1971 1971.3.16.1 | CAS 13396 | FMNH 70525 | ZMA 110740 Nematobrycon palmeri x |x x Eigenmann (1911) | Maldonado-Ocampo Eigenmann, 1911 et al. (2013b) Cheirodontops geayi Schultz, 1944 x Urbano-Bonilla et al. (2009). Waisted sw ansioni Steindachner (1880) | Eigenmann (1915) at Ne fakey. x X || IAVE-P 11746 | MCNG 24875, 24981, pa ger? 32355, 33285 | NMW 62543-44 es fe vanide amare Cone: x Fowler (1945) | Lima RS (2003) Ogee cepyg eubraheim X Urbano-Bonilla et al. (2009). & Malabarba, 2007 Odontostilbe pulchra (Gill, 1858) Odontostilbe splendida Bihrnheim & Malabarba, 2007 Biihrnheim and Malabarba (2007) Bihrnheim and Malabarba (2007) | ANSP 181040, 181041 | [AVH-P 3239, 3326, 3328, 3615, 7943, 7945, 7949, 7951, 8050, 8051, 8070, 9553-9555, 9557, 9558, 9697 | ICN-MHN 14168 | INPA 25173 | MBUCYV 32890 | MCNG 54519 | MCP 38862 54 Carlos DoNascimiento et al. / ZooKeys 708: 25-138 (2017) Taxa Protocheirodon pi (Vari, 1978) Saccoderma hastata DC EN | ET | Amz alee [ae a Ori Pac Car | References/Collections CAS 41749 (izentiahn, 1913) X | xX ible. 4 Eigenmann (1913) | Eigenmann (1915) Saccoderma melanostigma Schultz, 1944 X |Ortega-Lara et al. (2012) ie mpirasta Dalibs ™ 5h 5x X |Dahl (1955) | MHNG 1066.39-42 i Roman-Valencia (2000) | ICN-MNH ike Bie eRomnen 2722 | [AvH-P 4734 | IMCN 1689, ci 3894, 3905, 3911, 3918, 4052 Eretmobrycon emperador (Eigenmann & Osle, 1907) X_ |Maldonado-Ocampo et al. (2013b) Pe : Eigenmann et al. (1914) | FMNH 56657 ee toe oe ce pallies [ex CM 5474] | CAS 40844 [ex IU (Eigenmann & Henn, 1914) 13168] Eretmobrycon miraensis x |x Fowler (1945) | ANSP 71686, 71687-92, (Fowler, 1945) 71693, 71694 Eretmobrycon peruanus Maldonado-Ocampo et al. (2013b) | (Miller & Troschel, 1845) Fowler (1939) Eretmobrycon scleroparius Maldonado-Ocampo et al. (2006b), Co- (Regan, 1908) lombia. Biota Colombiana 7(1): 143-154 Markiana geayi (Pellegrin, 1909) Lasso et al. (2005) * Scopaeocharax atopodus (Bahlke, 1958) CIACOL 2713 Tyttocharax cochui , (Ladiges, 1950) Arbeldez et al. (2004) Tyttocharax metae Roman- Valencia, Garcfa-Alzate, X |X X Roman-Valencia et al. (2012) | [UQ 2581 Ruiz-C. & Taphorn B., 2012 Xenurobrycon heterodon x Bogota-Gregory and Maldonado-Ocampo Weitzman & Fink, 1985 (2006a) | USNM 280246 Chrysobrycon guahibo Vanegas-Rios, Urbano- xX .| X Xx Vanegas-Rios et al. (2015) Bonilla & Azpelicueta, 2015 PUL U EE Danes x Vanegas-Rios et al. (2013b) (Bohlke, 1958) Chrysobrycon mojicai Vanegas-Rios & Urbano- DG |X Vanegas-Rios et al. (2017) Bonilla, 2017 Corynopoma riisei Gill, 1858 Cala (1977) Eigenmann (1912) | CAS 44275-77 [ex Gephyrocharax caucanus x lx IU 12668-70] | FMNH 56012 [ex CM Eigenmann, 1912 4802], 56013-15, 69553, 95008 | USNM 81921 | CEP yracpanaecnacdenss mM | X X | X |Eigenmann (1912) | Vanegas-Rios (2016) Eigenmann, 1912 Checklist of the freshwater fishes of Colombia: a Darwin Core alternative... 55 . | Mag- ; Taxa DC |EN| ET | Amz| Ori Ge Pac| Car | References/Collections Gephyrocharax martae Dahl, x |x x Dahl (1943) 1943 Eigenmann (1912) | CAS 44292-93 [ex Gephyrocharax melanocheir x |x x x IU 12696-97] | FMNH 56049 [ex CM Eigenmann, 1912 4839], 56050-51, 69554 | USNM 79209 | Vanegas-Rios (2016) Gephyrocharax sinuensis x |x x Dahl and Medem (1964) | ICN-MHN Dahl, 1964 749 Gephyrocharax torresi Vanegas-Rios, Azpelicueta, ; Mies Ganncener ies XM.) X XxX Vanegas-Rios et al. (2013a) | UIST 1767 2013 Gephyrocharax valencia . Figentain, 1920 xX Vanegas-Rios (2016) Gephyrocharax venezuelae Schultz, 1944 X |Ortega-Lara et al. (2012) Pterobrycon landoni x lx Eigenmann (1913) | FMNH 56250 [ex Eigenmann, 1913 CM 5051] Lay Géry (1966b) | ANSP 111676 [ex Géry ee sr ates coll. 0124.18-19], 111668 | MNHN 1980-1425 | ZMA 113828 Hemibrycon antioquiae Roman Vane Ke) Roman-Valencia et al. (2013) Taphorn, Mancera-Rodriguez & Garcia-Alzate, 2013 Hemibrycon boquiae Eigenmann (1913) | CAS 44332 [ex IU (Roman-Valencia, 1999) xX xX xX ; xX | X xX 12831] | FMNH 56259 [ex CM 5059], cue ia SL eLE 56260 Hemibrycon brevispini Ronin alenciend Acie Men G009) Roman-Valencia & Arcila- X |X Xx | 1U0 5 4D ae asa maces Tb aca Mesa, 2009 Hemibrycon catroense , } : Roman-Valencia & Arcila- XxX. |X xX ‘moet and Pecil Meesak2009) Mesa, 2009 Hemibrycon cardalensis Roman-Valencia, Ruiz-C., Taphorn, Mancera- X |X Xx Roman-Valencia et al. (2013) Rodriguez & Garcia-Alzate, 2013 ee pincers | cdl x Dahl (1960c) | ICN-MHN 239, 31, 126 Hemibrycon caucanus x x Eigenmann (1913) | Maldonado-Ocampo (Eigenmann, 1913) et al. (2006b) Eigenmann et al. (1914) | MNH 56653 Hemibrycon colombianus x |x x [ex CM 5470], 56654-56 | CAS 44350 [ex Eigenmann, 1914 IU 13162], 44351 [ex IU 13163], 44352 [ex IU 13164], 44353 [ex IU 13165] Hemib nD: Roman-Valencia (1999) | IAVH-P 7529, a Es la ale Sele: 4 7530 | ICNMNH 3445, 3446-49 | IUQ 381-382 56 Carlos DoNascimiento et al. / ZooKeys 708: 25-138 (2017) . | Mag- ; Taxa DC |EN| ET | Amz| Ori ene Pac| Car| References/Collections Hemibrycon dariensis Meek : ' ; &¢ Hildebrand, 1916 las Roman-Valencia and Ruiz-C. (2007) Hemibrycon decurrens FMNH 56255 [ex CM 5055] | CAS (Eigenmann, 1913) 39542 [IU 12829] Hemibrycon dentatus : (Figenmann, 1913) S aale.€ Eigenmann (1913) Hemibrycon fasciatus Roman- Valencia, Ruiz-C., Taphorn, : ; iulsacersRcrisuezs Be x | Roman-Valencia et al. (2013) Garcia-Alzate, 2013 Hemibrycon galvisi (Roman- x |x Roman-Valencia (2000) | ICN-MHN Valencia, 2000) 2720, 2721 | TUQ 221-223 Hemibrycon jabonero Schultz, Ortega ara eral. 2012) 1944 Hemibrycon jelskii (Steindachner, 1877) X Maldonado-Ocampo et al. (2008) fie Diep CENCE x 3639, 7528, 9194-9197, 9203, 9677-9680 | MNHN 1980-1426 | USNM 198645 Hemibrycon metae Myers, om Pf) || poses a530 Hemibrycon microformaa } ; : Roman-Valencia and Ruiz-C. (2007) | Roman-Valencia & Ruiz-C., | X | X TUQ 510, 511-514 | MTDF 27628 2007 Hemibrycon paez Roman- Valencia & Arcila-Mesa, xX | X Xx Roméan-Valencia and Arcila-Mesa (2010) 2010 Hemibrycon palomae Roman- Romén-Valencia et al. (2010a) | [UQ Valencia, Garcfa-Alzate, K | X X 2727-2729, 2802, 2846, 2300, 2301, Ruiz-C. & Taphorn, 2010 AUM 50793 Hemibrycon plutarcoi x |x x Romén-Valencia (2001) | ICN-MHN (Roman-Valencia, 2001) 4887 | IUQ 308, 461, 472, 473 Hemibrycon quindos Roman- Valencia & Arcila-Mesa, Ks | OS Xx Roman-Valencia and Arcila-Mesa (2010) 2010 Hemibrycon rafaelense Roman-Valencia and Arcila-Mesa (2008) | Roméan-Valencia & Arcila- X |X xX ICN-MHN 6703, 3505 | IUQ 499, 509 | Mesa, 2008 MCNG 54101 | MTD F 27623-24 Hemibrycon raqueliae Roman-Valencia & Arcila- xX |X Roman-Valencia and Arcila-Mesa (2010) Mesa, 2010 Hemibrycon sanjuanensis Roman-Valencia, Ruiz-C., : . Tiphorise Canete Alvar xX |X xX Roman-Valencia et al. (2014b) 2014 Hemibrycon santamartae Roman-Valencia, Ruiz-C., : : GarclasAlzate 6 Tiphorn, X |X X_ |Roman-Valencia et al. (2010d) 2010 1913 Creagrutus machadoi Vari & Galvis et al. (2007a) Checklist of the freshwater fishes of Colombia: a Darwin Core alternative... 57 Taxa DC|EN|ET |Amz| Ori ae Pac| Car | References/Collections Hemibrycon sierraensis Garcia-Alzate, Roman- Oe |X X | Garcia-Alzate et al. (2015a) Valencia & Taphorn, 2015 ; Dahl and Medem (1964) | ICN-MHN Hemibrycon velox Dahl, 1964} X | X X xX 1457, 1463, 1464, 2711 Hemibrycon virolinica Roman-Valencia & Arcila- CM ie: x Roman-Valencia and Arcila-Mesa (2010) Mesa, 2010 Hemibrycon yacopiae Roman- Valencia & Arcila-Mesa, X |X Xx Roman-Valencia and Arcila-Mesa (2010) 2010 Carl sis Eigenmann (1913) | CAS 68647 [ex IU ae eed ie | exe xe) x 12911] | FMNH 56882 [ex CM 5162], ay 56883 Creagrutus affinis : : Steindachner, 1880 X | Steindachner (1880) | Eigenmann (1921) Creagrutus amoenus Fowler, Fowler (1943) | ANSP 70499, 70500-02, 1943 70503 Vari and Harold (2001) | ANSP 134080 | Creagrutus atratus Vari & x lx xl. IAvH-P 3659, 3660, 8426 | ICN-MHN Harold, 2001 16981 | NRM 16842-43 | USNM 353866-67, *Creagrutus barrigai Vari & Harold, 2001 xX CZUT-IC 12062 Creagrutus bolivari Schultz, x Vari and Harold (2001) 1944 Eigenmann (1913) | CAS 41341-42 [ex Creagrutus brevipinnis x |x x IU 12728-29], 41375 [ex IU 12730] | Eigenmann, 1913 FMNH 56095 [ex CM 4887a], 56096- 98, 75172 | USNM 79188 Vari and Harold (2001) | ANSP 130527, Creagrutus calai Vari & x lx xX 139149, 177718 | IAVH-P 7918 | NRM Harold, 2001 16848-49, 43015-16 | USNM 353304, 353307, 353868 1s oe Eigenmann (1913) | CAS 41373-74 [ex ‘ tks a. . Ae ly lei x IU 12736-37], 69304 [ex IU 12738] | Boo ae FMNH 56104 [ex CM 4895a], 56105-08 Creagrutus cochui Géry, 1964 xm | ey Vari and Harold (2001) Creagrutus flavescens Vari & Harold, 2001 X Vari and Harold (2001) C ; Tanne Torres-Mejia and Vari (2005) | ICN- Rien one iy - sal [>< x MHN 8520, 8521, 8522, 9893, 9894, 9895, 9896, 9897, 9898 Creagrutus hildebrandi Schultz, 1944 X | Ortega-Lara et al. (2012) Creagrutus leuciscus Regan, x |x x Regan (1913) | et yy Harold, 2001 58 Carlos DoNascimiento et al. / ZooKeys 708: 25-138 (2017) Taxa DC | EN| ET | Amz| Ori ae Pac| Car | References/Collections Creagrutus maculosus Roman-Valencia, Garcia- : Aleate Ruiz G.&'Taphem a |S Roman-Valencia et al. (2010b) B, 2010 Eigenmann (1913) | CAS 41641 [ex IU Creagrutus magdalenae x |x 12722], 60056-57 [ex IU 12725-26] | Eigenmann, 1913 FMNH 56088 [ex CM 4880], 56089, 56092-93, 69730 Creagrutus maracaiboensis Schultz, 1944 X | Ortega-Lara et al. (2012) Creagrutus maxillaris (Myers, Myers (1927) | [AVH-P 10192 1927) Creagrutus melasma Vati, MPUJ 8814, 8815, 9207, 9208, 9209 Harold & Taphorn, 1994 : : : ‘ Creagrutus nigrostigmatus x |x Dahl (1960b) | ICN-MHN 989 | SU Dahl, 1960 49491 Creagrutus paralacus Harold & Vari, 1994 X | Harold and Vari (1994) 1927 *Creagrutus runa Vari & Harold, 2001 x CZUT-IC 4117, 4163 Creagrutus taphorni Vari & Harold, 2001 x Urbano-Bonilla et al. (2009). Creagrutus tuyuka Vari & Lima, 2003 Creagrutus vexillapinnus Vari &¢ Harold, 2001 xX IAvH-P-14609, 14813 CN ee Ladle x | x x Fowler (1943) | ANSP 70505 (Fowler, 1943) *Ceratobranchia binghami Eigenmann 1927 * Ceratobranchia joanae Chernoff & Machado- Allison, 1990 ANSP 70504 Ceratobranchia obtusirustris Eigenmann, 1914 Knodus alpha (Eigenmann, 1914) x CZU TAG 12827 x Bogota-Gregory and Maldonado-Ocam- po (2006a) | [AVH-P 7526 Eigenmann et al. (1914) | CAS 18511 [ex IU 13157] | EBRG 295, 490, 493, 690, 822, 856, 886, 921, 922, 941, 1311 | FMNH 56646 [ex CM 5463], 56646-47 | [AVH-P 3558, 3640, 3641, 3643-3647, 3650-3653, 3657, 8042-8044, 8054, 8055, 9152-9158, 9178-9180, 9200- 9202, 9664-9676, 9781, 11789 Knodus breviceps (Eigenmann, 1908) IAvH-P 12289, 2538, 2575, 13165, 5176, 5187, 5418, 5419, 8315, 8316, 8317, 8318, 8319, 8320, 8321, 8322, 8323, 8324, 8325, 8326, 8327, 8328, 8329, 8330, 8655, 8694, 8729, 8764, 8795, 8828, 8858, 8889, 8920, 8942, 8954, 8974, 8990, 9013, 9026, 9047, 9072, 9097 Checklist of the freshwater fishes of Colombia: a Darwin Core alternative... 52 Taxa DC |EN| ET | Amz| Ori ae Pac| Car| References/Collections TD eo tipae OR natin IAvH-P-14385, 14396, 14443, 14610, Valencia. teshon By & xX 14637, 14705, 14806, 14811, 14994, : : : 15059, 15228, 15237, 15542, 15583, Ruiz-C., 2008 15603 Eigenmann et al. (1914) | FMNH 56644 eee & |x [ex CM 5461], 56645 | CAS 61221 [ex 2 TU 13156] *Knodus gamma Géry, 1972 | |x] o[ | | jezuric 12073, 12967 Knodus heteresthes x Roman-Valencia (2003b) | CZUT-IC (Eigenmann, 1908) 4151, 4200, 12308 cane ihenidae Bigenmann, X_ |Maldonado-Ocampo et al. (2008) pce Seager 4 x Arbeléez et al. (2004) | Bogota-Gregory mua ae sue and Maldonado-Ocampo (2006a) 1903) ie orteguasae (Fowler, x |x x Fowler (1943) Knodus tiquiensis Ferreira & xX Ferreira and Lima (2006) | CZUT-IC Lima, 2006 12307 *Rhinobrycon negrensis Argopleura chocoensis x |x x |x Eigenmann (1913) | Maldonado-Ocampo (Eigenmann, 1913) et al. (2006b) cop leita potions Eigenmann (1913) | CAS 39439 [ex IU ete ie) x |x Xx 12802] | FMNH 56261 [ex CM 5060], ; 56262, 71290 | USNM 79174; Aaah eave daa Eigenmann (1913) | CAS 39013 [ex IU ane er x |x 12820] | FMNH 56272 [ex CM 5072], : 56273, 69690; Eigenmann (1913) | BMNH “Arsuplouta mnbaalonenie 1924.3.3.37-38 | CAS 40827-32 [ex (Eisenman, 1913) X |X IU 12821-26] | FMNH 56263 [ex CM , 5063], 56264-65, 56267-71, 56275, 69760 | USNM 79176, 236097 We CZUTAC 18065, MPUJ 273, 1257, Marinho, 2013 9846-9847, 10726-10729, 10744 Othonocheirodus Myers, CZUT-IC 5497, 12031, 12033, 12525, 1927 MPUJ 3747 Bove eidtharibue eRe! Roman-Valencia (2003a) | IUQ 447, Rowse Malenets 2003 x |X 448-449 | CAS 70099-101 | FMNH ; 56566-67 Bryconamericus arilepis ; : Roman-Valencia, Vanegas- X | X Xx ne aera al ODE? NO 2205 Rios & Ruiz-C, 2008 ; , Bryconamericus caldasi Roman-Valencia, Ruiz-C., x |x x Roman-Valencia et al. (2014a) | I[UQ Taphorn B. & Garcia-Alzate, 2014 3714 Bryconamericus carlosi Roman-Valencia, 2003 Roman-Valencia (2003c) 60 Carlos DoNascimiento et al. / ZooKeys 708: 25-138 (2017) T . | Mag- ‘ axa DC |EN| ET | Amz| Ori ene Pac| Car| References/Collections Roman-Valencia (2003c) | FMNH 56642 Bryconamericus cismontanus x [ex CM 5459] | [AVH-P 3315, 3317- Eigenmann, 1914 3319, 7920, 9168-9170, 9188-9193, 9480, 9481, 9650-9663, 9753 Bryconamericus foncensis Roman-Valencia, Vanegas- X |X Xx Roman-Valencia et al. (2009) | [UQ 1941 Rios & Ruiz-C, 2009 Bryconamericus guizae x |x ily Romén-Valencia (2003a) | TUQ 450, Roman-Valencia, 2003 451-452 | STRI 1370 Berne nen ns niilae Roman-Valencia (2003a) | IUQ 422, Romén-Valencia, 2003 xX |X xX 312-313, 423-424, 462-463, 476-477) | : STRI 00574 : et rrmparae Galles aa lo X |Dahl and Medem (1964) | SME 16284 Bryconamericus ichoensis x |x x Roman-Valencia (2000) | ICN-MHN Roman-Valencia, 2000 2718, 2719 Bee ee oe, Roméan-Valencia et al. (2010c) | [UQ ae Ree & aephor X | X xX 2448, 2271, 2326, 2435, 2437, 2440- ° ; . 2447, 2559-2561 | AUM 50297 2010 *Bryconamericus macrophthalmus Roman- X CZUT-IC 5124 Valencia, 2003 panic mined x xP TE || | x foatto rex aa *Bryconamericus orinocoense Roman-Valencia, 2003 = CAUTAC ABO A216 Sete en a |e x Eigenmann (1913) Mictovenysteinger Eigenmann (1913) | CAS 47170 [ex IU ; X01] EX, | EX xX 12818] | FMNH 56215 [ex CM 5007], Eigenmann, 1913 56216 Astyanacinus yariguies Torres-Mejia, Hernandez & | X | X X Torres-Mejia et al. (2012) | UIST 1752 Senechal, 2012 ia hog go Wes! at IAVH-P 8238-8240, 9117, 11012 Eigenmann, 1908 Eee ate > ane. ¢ X |Eigenmann and Ogle (1907) Avsindn biradoulana Fowler (1943) | Maldonado-Ocampo (Linnaeus, 1758) X {et al. (2013b) | Steindachner (1879a) | : IAvH-P 1694, 9110 Astyanax caucanus x |x Steindachner (1879c) | NMW 57372-76 | (Steindachner, 1879) SMNS 2833 | ZMUC 993 Astyanax daguae Eigenmann, x|xlx Eigenmann (1913) | Maldonado-Ocampo 1913 et al. (2006b) Astyanax fasciatus Eigenmann (1922) | Fowler (1943) | Lima (Cuvier, 1819) et al. (2003) | Eigenmann (1921) Astyanax fasslii : a a pee iS) x2] al Steindachner (1915) Eigenmann, 1912 Jupiaba abramoides 4808], 56019-2, 69547, 69779 | USNM 79207 Cala (1977) | ICN-MHN 546 Checklist of the freshwater fishes of Colombia: a Darwin Core alternative... 61 Taxa DC | EN| ET | Amz| Ori ae Pac| Car| References/Collections Astyanax filiferus x ae 4 Eigenmann (1913) | Maldonado-Ocampo (Eigenmann, 1913) et al. (2006b) Astyanax gisleni Dahl, 1943 | X | X Se al Dahl (1943) *Astyanax guianensis ; ea cat 1909 ‘ Maninbe (2015) Myers (1930) | [AVH-P 3300, 3305-3307, 23726 Astyanax magdalenae x Eigenmann and Henn (1916) | Maldona- Eigenmann & Henn, 1916 do-Ocampo et al. (2006b) Astyanax maximus Sea (le Bogota-Gregory and Maldonado-Ocam- (Steindachner, 1876) po (2006a) | [AvH-P 1731 Astyanax megaspilura Fowler, x |x | [| pees Fowler (1944) | Maldonado-Ocampo et 1944 al. (2006b) Eigenmann et al. (1914) | CAS 39229 Astyanax metae Eigenmann, [ex IU 13153] | CRPUT 302, 305, 527, 1914 x 331, 332 | FMNH 56640 [ex CM 5457], 56641 | IAVH-P 3302, 3521, 3522, 3525- 3531, 3535-3544, 7909, 9484, 9643-964 | Eigenmann (1913) | CAS 39341-45 [ex Astyanax microlepis x |x IU 12769-73] | FMNH 56209 [ex CM Eigenmann, 1913 5001], 56210-14, 69559-61, 71291, 95006 | USNM 79167 Astyanax orthodus x |x ESE: Eigenmann and Ogle (1907) | Maldona- Eigenmann, 1907 do-Ocampo et al. (2006b) Astyanax ruberrimus lx Eigenmann (1913) | Maldonado-Ocampo Eigenmann, 1913 et al. (2006b) | Eigenmann (1921) Astyanax scintillans Myers, x IAvEL_P 4979 1928 Astyanax siapae Garutti, Astyanax stilbe (Cope, 1870) SS) seal Eigenmann (1921) ple at ca at xX Urbano-Bonilla et al. (2009). 1944 ae Si Xx Weitzman et al. (2005) EC. T. Lima (pers. com.): as Hemi- Bryconella pallidifrons x grammus analis in Galvis et al. (2006) | (Fowler 1946) CZUT-IC 17948, IAvH-P 2605, MPUJ 12206, 12207 *Ctenobrycon oliverai Benine, x CZUTIC T2175. 9056;9279;-9321, Lopes & Ron, 2010 9670, 9752, 9785, 9801, 9810, 9851 ie fe “Vesely 7 Mojica et al. (2005) | Galvis et al. (2007a) Eigenmann (1912) | CAS 44271-73 [ex Genycharax tarpon x lxlx xX IU 12672-74] | FMNH 56018 [ex CM tt JH (Eigenmann, 1909) 62 Carlos DoNascimiento et al. / ZooKeys 708: 25-138 (2017) . | Mag- ; Taxa DC|EN|ET | Amz| Ori Gi Pac| Car| References/Collections Jupiaba anteroides (Géry, x |x Arbelaez et al. (2004) | Galvis et al. 1965) (2007a) Jupiaba asymmetrica (Eicennhaim 1908) X Fowler (1945) | Lima et al. (2003) Jupiaba polylepis (Ginther, x TAvELP 2013 1864) Jupiaba scologaster (Weitzman & Vari, 1986) a Cabs etal (20070) Jupiaba zonata (Eigenmann, x Galvis et al. (2007b) 1908) oe ae Schultz (1966) | USNM 231738 [was bate i566 J KC al'S 4 USNM 257403-F27], 171751, 231739 : [ex USNM 258173-F1] ite Géty (1972) | MHNG 1226.90., Pseudochalceus longianalis : Géry, 1972 1226.91-99 | ANSP 140067 | Géry Schultzites axelrodi Géry, x 1964 Serrabrycon magoi Vari, 1986 Thrissobrycon pectinifer Bohlke, 1953 ss personal collection 0690 x x Géry (1964a) | USNM 198642 xX FMNH 92642 Xx Maldonado-Ocampo et al. (2006a) Gasteropelecidae 9 spp oo marthae Myers, Ey | Se Mojjica et al. (2005) | Cala (1977) Cormegally payers: Fetnarider- x Mojica (1999) | NRM 200 | IMCN 7062 Yépez, 1950 Carnegiella schereri Fernandez-Yépez, 1950 Carnegiella strigata (Giinther, 1864) Galvis et al. (2007b) Weitzman (1960) | Cala (1977) Engraulisoma taeniatum Castro, 1981 Gasteropelecus maculatus Steindachner, 1879 Gasteropelecus sternicla (Linnaeus, 1758) TAvH-P 13957 Eigenmann (1912) | Eigenmann (1922) | Weitzman (1960) Weitzman (1960) Thoracocharax securis (De Filippi, 1853) Mojica et al. (2005) Thoracocharax stellatus (Kner, 1858) Bryconidae 22 spp (2 added and 3 deleted from 2008) Weitzman (1960) Brycon amazonicus (Agassiz, 1829) Brycon argenteus Meek & Hildebrand, 1913 Brycon dentex Giinther, 1860 Maldonado-Ocampo et al. (2006a) | TAvH-P 11603 Maldonado-Ocampo et al. (2013b) Mojica et al. (2004) | ICN-MHN 2351, 5025 Checklist of the freshwater fishes of Colombia: a Darwin Core alternative... 63 Steindachner, 1880 MHNG 1226.90, 1226.91-99 Salminus hilarii Fowler (1943) | Cala (1977) Taxa DC|EN|ET|Amz|O trae | Pac|Car| References/Collections Bogota-Gregory and Maldonado- Brycon falcatus Miiller & Ocampo (2006a) | Maldonado-Ocampo Troschel, 1844 et al. (2006a) | [AVH-P 5326 | ICN- MHN 4260, 2000, 5677 Brycon fowleri Dahl, 1955 X X |Dahl (1955) | ICN-MHN 2044 Brycon henni Eigenmann, x Eigenmann (1913) | Maldonado-Ocampo 1913 et al. (2005) *Brycon hilarii Brycon labiatus Steindachner, 1879 X |X Steindachner (1879b) | NMW Brycon medemi Dahl, 1960 x X |Dahl (1960c) | ICN-MHN 41, 73 Brycon meeki Eigenmann & x Eigenmann (1918a) | Maldonado- Hildebrand, 1918 Ocampo et al. (2006b) on melanopterus (Cope, CECT Mojica et al. (2005) | Galvis et al. (2007a) Brycon moorei x |x Steindachner (1878) | Mojica et al. Steindachner, 1878 * | (2006b) rae, Regan (1913) | Maldonado-Ocampo et al. Brycon oligolepis Regan, 1913 (2006b) | Eigenmann (1922) Brycon pesu Miller & Troschel, 1845 Galvis et al. (2007b) | Galvis et al. (2007a) *Brycon polylepis Moscé Morales 1988 Lima (2017) | [AVH-P 3554, 3671 Brycon posadae Fowler, 1945 Fowler (1945) | ANSP 71695 Brycon rubricauda Steindachner, 1879 |X| ES Steindachner (1879b) Brycon sinuensis Dahl, 1955 x |X X | Dahl (1955) Brycon striatulus (Kner, Regan (1913) | Maldonado-Ocampo et 1863) al. (2008) | Brycon whitei Myers & x |x Myers and Weitzman (1960) | Urbano- Weitzman, 1960 Bonilla et al. (2009). Salminus affinis Steindachner (1880) | ANSP 140067 | Valenciennes, 1850 Triportheidae 12 spp (2 added and 2 deleted from 2008) Agoniates anchovia Eigenmann, 1914 Mojica et al. (2005) Agoniates halecinus Miiller & Troschel, 1845 Triportheus albus Cope, 1872 Bogota-Gregory and Maldonado- Ocampo (2005) | Maldonado-Ocampo et al. (2006a) Mojica et al. (2005) Triportheus angulatus (Spix & Agassiz, 1829) Mojica et al. (2005) Triportheus auritus (Valenciennes, 1850) Maldonado-Ocampo and Bogotd-Gregory (2007) | Galvis et al. (2007b) 64 Carlos DoNascimiento et al. / ZooKeys 708: 25-138 (2017) Tava DC|EN|ET | Amz| Ori pd Pac| Car| References/Collections es fe] | | feats rn Eee culter (Cope, X CZUT-IC 7310 Triportheus magdalenae xX X Steindachner (1878) | NMW 69151-54 | (Steindachner, 1878) ZMUC 87 nea ee x Maldonado-Ocampo et al. (2013a) ae pictus (Garman, xX Correa (2003) as EOE X IAvH-P 11059 ees re ad x Galvis et al. (2007a) Iguanodectidae 12 spp (1 added and 1 deleted from 2008) Bryconops affinis (Giinther, 1864) Bogota-Gregory and Maldonado-Ocampo (2005) | Cala (1977) | IAVH-P 2725, 3579 Bryconops alburnoides Kner, 1858 Bryconops caudomaculatus (Giinther, 1864) Galvis et al. (2007b) | ICN-MHN 9918 Correa (2003) | Eigenmann (1922) Bryconops collettei Chernoff & Machado-Allison, 2005 Maldonado-Ocampo et al. (2008) Bryconops humeralis Machado-Allison, Chernoff & Buckup, 1996 Maldonado-Ocampo et al. (2006a) Chalceus epakros Zanata & i Zanata and Toledo-Piza (2004) | Bogota-Gregory and Maldonado- Bryconops giacopinii x |x Ocampo (2006a) | Maldonado-Ocampo (Ferndndez-Yépez, 1950) and Bogoté-Gregory (2007) | [AvH-P 1691, 5307 | ICN-MHN 4625, 10371 Bryconops inpai Knéppel, x |x Mojica et al. (2005) | Maldonado- Junk & Géry, 1968 Ocampo et al. (2006a) *Bryconops magoi Chernoff & Machado-Allison, 2005 a CAUPIC a2? se adujai Gécy, | laa Maldonado-Ocampo et al. (2008) Iguanodectes geisleri Géry, x |x Castro and Arboleda (1988) | Maldonado- 1970 Ocampo (2001) Iguanodectes purusii bs (Steindachner, 1908) = iMoneuct al 20) Iguanodectes spilurus x |x Arbeldez et al. (2004) | Maldonado- (Giinther, 1864) Ocampo et al. (2006a) Chalceidae 3 spp Toledo-Piza, 2004 Chalceus erythrurus (Cope, 1870) x Maldonado-Ocampo et al. (2006a) Mojica et al. (2005) Checklist of the freshwater fishes of Colombia: a Darwin Core alternative... 65 Taxa DC | EN| ET | Amz| Ori ae Pac| Car | References/Collections Chalceus macrolepidotus Cuvier 1816 Correa (2003) | Cala (1977) Gymnotiformes 78 spp (12 added and 8 deleted from 2008) Gymnotidae 12 spp (2 deleted from 2008) Electrophorus electricus Maldonado-Ocampo and Albert. (2003) (Linnaeus, 1766) | Cala (1977) Gymnotus anguillaris : Hocdeman, 1963 Maldonado-Ocampo and Albert. (2003) ere ane Maldonado-Ocampo and Albert (2004) | Maldonado-Ocampo & XxX IAvH_P 3477, 4001 Albert, 2004 “i : eae PEP OTINNA CES; Fowler (1945) | Eigenmann (1922) hi PPS CREEP ARO: Maldonado-Ocampo and Albert (2003) Leccia, 1994 Gymnotus choco Albert, ‘ Chnintaerneldesade, We Albert and Crampton (2003b) | ICN Ocampo, 2003 MHN 6621 | NRM 27734 Gymnotus coropinae Hoedeman, 1962 Gymnotus henni Albert, Crampton & Maldonado- Ocampo, 2003 Maldonado-Ocampo and Albert (2003) | Galvis et al. (2007a) Albert and Crampton (2003b) | CAS 47290, 217162 | EMNH 56793 Gymnotus javari Albert, Crampton & Hagedorn, 2003 Gymnotus pedanopterus Mago-Leccia, 1994 Gymnotus stenoleucus Mago- Galvis et al. (2007b) Maldonado-Ocampo and Albert (2003) Leccia, 1994 x | aX Maldonado-Ocampo and Albert (2003) Gymnotus tigre Albert & X Albert and Crampton (2003b) | UF Crampton, 2003 25552, 128412 | ICN-MHN 6690 Hypopomidae 9 spp (5 added and 2 deleted from 2008) Brachyhypopomus batesi Crampton, de Santana, Waddell & Lovejoy, 2016 Brachyhypopomus beebei (Schultz, 1944) Brachyhypopomus bennetti Sullivan, Zuanon & Cox Fernandes, 2013 Crampton et al. (2016) Maldonado-Ocampo and Albert (2003) Sullivan et al. (2013b) Brachyhypopomus brevirostris (Steindachner, 1868) Brachyhypopomus bullocki Sullivan & Hopkins, 2009 Maldonado-Ocampo and Albert (2003) Sullivan and Hopkins (2009) | Crampton et al. (2016) | ANSP 187477 66 Carlos DoNascimiento et al. / ZooKeys 708: 25-138 (2017) Taxa DC|EN| ET |Amz| Ori ae Pac| Car| References/Collections Brachyhypopomus flavipomus Crampton, de Santana, Waddell & Lovejoy, 2016 Crampton et al. (2016) Brachyhypopomus occidentalis (Regan, 1914) Regan (1914) | Maldonado-Ocampo et al. (2006b) | Albert and Crampton (2003a) Brachyhypopomus sullivani Crampton, de Santana, Waddell & Lovejoy, 2016 Microsternarchus bilineatus Fernandez-Yépez, 1968 Crampton et al. (2016) Maldonado-Ocampo and Albert (2003) Rhamphichthyidae 10 spp (1 added and 1 deleted from 2008) Gymnorhamphichthys hypostomus Ellis, 1912 Maldonado-Ocampo and Albert (2003) Gymnorhamphichthys rondoni (Miranda Ribeiro, 1920) Maldonado-Ocampo and Albert (2003) *Rhamphichthys apurensis (Fernandez-Yépez, 1968) Rhamphichthys drepanium Triques, 1999 Rhamphichthys marmoratus Castelnau, 1855 CZUTF-IC 3244, 5178, 5179, 9362, 9421, 9422, 9423, 9424, 9448, 9449, 9547 Maldonado-Ocampo and Albert (2003) Cala (1977) Rhamphichthys rostratus (Linnaeus, 1766) Hypopygus lepturus Hoedeman, 1962 Hypopygus neblinae Mago- Leccia, 1994 Maldonado-Ocampo and Albert (2003) Maldonado-Ocampo and Albert (2003) Galvis et al. (2007b) | Maldonado- Ocampo and Albert (2003) Steatogenys duidae (La Monte, 1929) Maldonado-Ocampo et al. (2006a) | Maldonado-Ocampo and Albert (2003) Steatogenys elegans (Steindachner, 1880) Sternopygidae 16 spp (2 added from 2008) Maldonado-Ocampo and Albert (2003) Distocyclus conirostris (Eigenmann & Allen, 1942) ‘Eigenmannia goajira Schultz, 1949 Eigenmannia humboldtii (Steindachner, 1878) act Maldonado-Ocampo and Albert (2003) Maldonado-Ocampo and Albert (2003) Steindachner (1878) | Maldonado- Slows et al. (200Gb) Eigenmannia limbata (Schreiner & Ribeiro, 1903) Maldonado-Ocampo and Albert (2003) Eigenmannia macrops (Boulenger, 1897) Eigenmannia nigra Mago- Leccia, 1994 Eigenmannia virescens (Valenciennes, 1836) Maldonado-Ocampo and Albert (2003) Maldonado-Ocampo and Albert (2003) Eigenmann and Ward (1905) | Maldonado-Ocampo and Albert (2003) Checklist of the freshwater fishes of Colombia: a Darwin Core alternative... 67 . | Mag- ' Taxa DC|EN| ET | Amz| Ori Ge Pac| Car| References/Collections Rhabdolichops caviceps (Ferndndez- Yéper, 1968) Jeri Maldonado-Ocampo and Albert (2003) *Rhabdolichops eastwardi Lundberg & Mago-Leccia, X CZUT-IC 3642, 3987 1986 Rhabdolichops troscheli (Kaup, 1856) xe Maldonado-Ocampo and Albert (2003) *Rhabdolichops zareti Lundberg & Mago-Leccia, CZUT-IC 9471 1986 Sternopygus aequilabiatus Humboldt (1805) | Maldonado-Ocampo (Humboldt, 1805) et al. (2013b) weuee DSTI NRED X Maldonado-Ocampo and Albert (2003) Leccia, 1994 Sternopygus dariensis Meek & Hildebrand, 1916 | lee wae Sternopygus macrurus (Bloch x |x Maldonado-Ocampo and Albert (2003) | & Schneider, 1801) Eigenmann (1922) Sternopygus pejeraton Schultz, X |Ortega-Lara et al. (2012) 1949 Apteronotidae 31 spp (4 added and 3 deleted from 2008) Adontosternarchus balaenops (Cope, 1878) x Maldonado-Ocampo and Albert (2003) Adontosternarchus clarkae Mago-Leccia, Lundberg & x Mago-Leccia et al. (1985) Baskin, 1985 Adontosternarchus devenanzii Mago-Leccia, Lundberg & x Mago-Leccia et al. (1985) Baskin, 1985 *Adontosternarchus sachsi x CZUT-IC 9438, 9451, 9453, 9455, (Peters, 1877) 9457, 9475, 9490, 9492 Apteronotus albifrons (Linhaslis 1760) x Maldonado-Ocampo and Albert (2003) Apteronotus anu de Santana & Vari, 2013 X |de Santana and Vari (2013) Apteronotus apurensis Fernéndez-Yépez, 1968 ea ig Maldonado-Ocampo and Albert (2003) Apteronotus bonapartii (Castelnau, 1855) x Maldonado-Ocampo and Albert (2003) Apteronotus cuchillejo (Schultz, 1949) X_ |Maldonado-Ocampo and Albert (2003) ee aici seal, X_ |Maldonado-Ocampo and Albert (2003) Apteronotus eschmeyeri de Santana, Maldonado- x |x x de Santana et al. (2004) | Maldonado- Ocampo, Severi & Mendez, 2004 Ocampo et al. (2006b) 68 Carlos DoNascimiento et al. / ZooKeys 708: 25-138 (2017) T . | Mag- ‘ axa DC |EN| ET | Amz| Ori ene Pac| Car| References/Collections Apteronotus galvisi de Santana, Maldonado- Oe |G x de Santana et al. (2007) Ocampo & Crampton, 2007 RAIL AA eee x |X X | | Fowler (1944) | ANSP 71435 Apteronotus macrolepis x Bogota-Gregory and Maldonado- (Steindachner, 1881) Ocampo 2006 aa Gee eta x |X CBT Fowler (1943) | ANSP 70528 Apteronotus magdalenensis xlxlx x Miles (1945) | BMNH 1947.7.1.138 | (Miles, 1945) USNM 123795 Wpisroncms rice Eigenmann and Fisher (1914) | FMNH (Eigenmann & Fisher, 1914) es x 56774 [ex CM 5594] | CAS 62345 [ex ; IU 13375] de Santana and Maldonado-Ocampo. Apteronotus milesi de Santana (2005) | [AVH-P 3996 | CAS 72249 [ex IU & Maldonado-Ocampo, Xgl eX: x 13379], 72250 [ex IU 13379] | IAvH-P 2005 3997 | ICMN 2500 | ICNMNH 3155 | FMNH 56775, 56776, 56777, 56778 Apteronotus rostratus (Meek de Santana and Vari (2013) | Eigenmann & Hildebrand, 1913) and Fisher (1914) spp reranapes spiel Began) | 5 Regan (1914) | BMNH 1914.5.18.90-93 1914) Parapteronotus hasemani (Ellis, 1913) Platyurosternarchus macrostomus (Giinther, 1870) Maldonado-Ocampo and Albert (2003) Maldonado-Ocampo and Albert (2003) Sternarchorhynchus mormyrus (Steindachner, 1868) Sternarchorhynchus oxyrhynchus (Miiller & Troschel, 1848) Maldonado-Ocampo and Albert (2003) Maldonado-Ocampo and Albert (2003) Sternarchorhynchus roseni Mago-Leccia, 1994 Maldonado-Ocampo and Albert (2003) | IAvH-P 2981 Compsaraia compsus (Mago- Leccia, 1994) Sternarchogiton nattereri (Steindachner, 1868) *Sternarchella orinoco Mago- Leccia, 1994 Maldonado-Ocampo and Albert (2003) Maldonado-Ocampo and Albert (2003) CZUT-IC 9493, 9535 * Sternarchella orthos Mago- Leccia, 1994 Sternarchella schotti (Steindachner, 1868) Sternarchorhamphus muelleri (Steindachner, 1881) CZUT-IC 9445, 9482, 9484 Maldonado-Ocampo and Albert (2003) | Evans et al. (2017) Maldonado-Ocampo and Albert (2003) Siluriformes 588 spp (98 added and 61 deleted from 2008) Checklist of the freshwater fishes of Colombia: a Darwin Core alternative... 69 T . | Mag- ; axa DC|EN| ET | Amz| Ori Ge Pac| Car|References/Collections Trichomycteridae 66 spp (16 added and 6 deleted from 2008) me aren gic | Se Xx Humboldt (1805) Ttuglanis guayaberensis (Dahl, x |x x Dahl (1960a) 1960) Ttuglanis metae (Eigenmann, ICe Tes Eigenmann (1917a) | CAS 58138 [ex IU 1917) 13770] Rhizosomichthys totae (Miles, x|xlx : ; Miles (1942) | ICN-MHN 353 | MCZ 1942) 35744 | SU 37074 Trichomycterus arhuaco Ardila x |x x Ardila Rodriguez (2016c) | Holotype Rodriguez, 2016 CAR 684 one idk i eet Se ee i LL Sie Ardila Rodriguez (201 1a) | CAR 400 Eigenmann (1912) | CAS 58127 Trichomycterus banneaui x |x x [ex IU 12677] | Holotype FMNH (Eigenmann, 1912) 56025 [ex CM 4815], 56026, 69815 | USNM 79234 Eigenmann (1912) | CAS 58118 [ex IU Trichomycterus bogotensis x |x x 12679] | Holotype FMNH 56030 [ex (Eigenmann, 1912) CM 4820] | 56031, 56044 | AMNH 7107 | USNM 79232 (EAN ae Ardila Rodriguez (2008c) | CAR 125, Ardila Rodriguez, 2008 X.| X | X xX 97 | CZUT-IC2920 | IAVH-P 11114 | : MBUCV-V35384; Trichomycterus caliensis x|xlx x |- Eigenmann (1912) | FMNH 56029 [ex (Eigenmann, 1912) CM 4819 not 6819} dene | OX X |Ardila Rodriguez (2017b) Eigenmann (1912) | CAS 58128 [ex Trichomycterus chapmani x |x calles IU 12678] | Holotype FMNH 56027 (Eigenmann, 1912) [ex CM 4817], 50628, 69813 | USNM 7235 i Fishopticieiaaiaaan Eigenmann (1917a) | CAS 64579 [ex IU (Figenmann, 1917) X | X xX 13810] | Holotype FMNH 58096 [ex 7 CM 7093], 58097 * Trichomycterus emanueli (Schultz, 1944) ial ei laid Trichomycterus : ; garciamarquezi Ardila x | xX a a COT6sh Hglotpe Rodriguez, 2016 Trichomycterus gorgona x|xlx x Fernandez and Schaefer (2005) | ANSP Fernandez & Schaefer, 2005 149946 | ICN-MHN 10019 Trichomycterus kankuamo x |x x Ardila Rodriguez (2016c) | Holotype Ardila Rodriguez, 2016 CAR 685 Trichomycterus knerii Stdindachnes, 1882 z Pa deh es Trichomycterus latidens x |x lk hese Eigenmann (1917a) | CAS 76335 [ex IU (Eigenmann, 1917) 13801] 70 Carlos DoNascimiento et al. / ZooKeys 708: 25-138 (2017) EN| ET | Amz ae P (Eigenmann, 1917) Trichomycterus striatus (Meek & Hildebrand, 1913) + |Eigenmann (1918b) Trichomycterus taenia Kner, XxX Regan (1913) Taxa DC Ori Cau |Pae Car| References/Collections Trichomycterus latistriatus x |x soll eel esr Lon ed Eigenmann (1917a) | FMNH 58449 (Eigenmann, 1917) [ex CM 7450] Trichomycterus maldonadoi x |x x Ardila Rodriguez (2011b) | CAR 380, Ardila Rodriguez, 2011 500 Trichomycterus manaurensis x |x x Ardila Rodriguez (2016c) | Holotype Ardila Rodriguez, 2016 CAR 683 lence oes x |omega-tara eal. 2012) HE hes ae Tee x |X x Dahl (1960a) | ICN-MHN 399, 400 Trichomycterus montesi Ardila x |x x Ardila Rodriguez (2016c) | Holotype Rodriguez, 2016 CAR 681 Scan etal Ss X | Ardila Rodriguer (2014) al Boulenger, x, | X - - Bourengen(i667) | BbANt 1887 1880.2.26.16-17 Trichomycterus ocanaensis x |x Ardila Rodriguez (2011d) | [AvVH-P Ardila Rodriguez, 2011 11115, 9797 Trichomycterus regani x |x x Eigenmann (1917a) | CAS 64591 [ex IU (Eigenmann, 1917) 13772] LCT BE REPRESS || Se X Regan (1903) | BMNH 1899.8.21.12-13 Regan, 1903 LRG oE x |x Xx Fowler (1941) | ANSP 69331, 69332-35 (Fowler, 1941) ie Momene na oaitdeeriess Ardila Rodriguez (2007) | CAR 340, 37, Ardila-Rodriguez, 2007 me |) OS x 88, 89, 325, 331, 332 | [AVH-P 4342, 4344 | IMCN 4195 rae cin se SU iprelexs x Ardila Rodriguez (2006a) | CAR 116 Trichomycterus santanderensis x |x x Castellanos-Morales (2007) | CAC- Castellanos-Morales, 2007 CDMB 035 Trichomycterus sketi x |x x Castellanos-Morales (2011) | CAC- Castellanos-Morales, 2011 CDMB 104 Trichomycterus spilosoma x |x ix Regan (1913) | BMNH 1910.7.11.106- (Regan, 1913) 107, 1910.7.11.108 Trichomycterus steindachneri LEN Ge aaere saa x DoNascimiento et al. (2014) Pedreros & Guerrero- Kommnritz, 2014 hemetee wean Eigenmann (1918b) | CAS 58121 [ex IU (Eienmanasto1s) oe ROS xX 13814] | FMNH 58101 [ex CM 7097], 58102 chahapetarad sedate Eigenmann (1917a) | CAS 58148 [ex IU X |X x - |13818], 58105 [ex IU 13804] | FMNH : 58105 [ex CM 7101], 58091-92 1863 Checklist of the freshwater fishes of Colombia: a Darwin Core alternative... Fal . | Mag- ‘ Taxa DC/|EN| ET | Amz| Ori Ge Pac| Car| References/Collections Trichomycterus tetuanensis Garcia-Melo, Villa-Navarro | X | X X Garcia-Melo et al. (2016) & DoNascimiento, 2016 Trichomycterus torcoromaensis . ¥ Ardila-Rodriguez, 2016 ou OG X Ardila Rodriguez (2016a) Trichomycterus transandianus (Steindachner, 1915) | SX: x Steindachner (1915) | NMW 44475 Trichomycterus uisae x |x x Castellanos-Morales (2008) | CAC- Castellanos-Morales, 2008 CDMB 072 Trichomycterus unicolor (Regan, 1913) X | X xX Regan (1913) | BMNH 1913.10.1.42-43 Malacoglanis gelatinosus x |x x | Myers and Weitzman (1966) | CZUT-IC Myers & Weitzman, 1966 13816, [AVH-P 13640, SU 50754, 50755 Tridens Eigenmann & x ICN-MHN 10101, 10330, 12112, Eigenmann, 1889 ‘ 12114 Tridensimilis venezuelae Schultz, 1944 : X |Ortega-Lara et al. (2012) ag pearsoni Myers, X CZUTC 17989, 18069 Haemomaster venezuelae Myers, 1927 Xx Maldonado-Ocampo et al. (2006a) Henonemus punctatus s (Boulenger, 1887) se Mojica et al. (2005) Henonemus triacanthopomus DoNascimiento & X Lasso et al. (2009) | CZUT-IC 9648 Provenzano, 2006 Megalocentor echthrus de Ochmacanthus alternus aioe x Dahl (1960a) Ochmacanthus orinoco Myers, x IAVH-P 14250-14252 1927 Ochmacanthus reinhardtii ’ (Steindachner, 1882) x Arbeldez et al. (2004) Pseudostegophilus haemomyzon (Myers, 1942) Bvid Fe 127, 83/0997 Pseudostegophilus nemurus Mojica et al. (2005) | [AVH-P 1189, 4996 (Giinther, 1869) rele i Schultzichthys bondi (Myers, MaldnideO citnporer al: (2008) 1942) Stegophilus septentrionalis Myers, 1927 xX CZUT-IC 5049 Paracanthopoma Giltay, 1935 x Maldonado-Ocampo et al. (2008) Paravandellia phaneronema x lx xX Miles (1943) | MCZ 35874 | USNM (Miles, 1943) 120141 Vandellia beccarii Di Gaporiaces, 1935 oll pl be | Maldonado-Ocampo et al. (2013a) 72 Carlos DoNascimiento et al. / ZooKeys 708: 25-138 (2017) Taxa DC|EN|ET|Amz|O ieee "| Pac| Car| References/Collections Vandellia cirrhosa Castro and Arboleda (1988) | de Pinna Valenciennes, 1846 and Wosiacki (2003) Callichthyidae 44 spp (2 added and 3 deleted from 2008) Mojica et al. (2005) | Cala (1977) | [AvVH- eu eat P 2089, 12161 | ICN-MHN 548, 6377, i 7494, 7495, 16897 : _— } Roméan-Valencia et al. (1999) | ICN- ers ellie oe . MHN 3842 | IUQ 152, 305, 306, 307 cicadas | MCP 21174-75 | UV 89018, 91063, eg 93001-04, 98002, 98031 Ardila Rodriguez (2006b) | CAR 251, Callichthys oibaensis Ardila- x 250 | IMCN 3310 | CZUT-IC 1837 Rodriguez, 2006 | [AvH-P 5730 | ICNMNH 13396 | MBUCYV-V 32798 Dianema longibarbis Cope, ‘Heectr Mojica et al. (2005) 1872 Hoplosternum littorale : (Hancock, 1828) Mojica et al. (2005) | Cala (1977) Hoplosternum magdalenae ' Ricentaaunt 1913 X_ |Eigenmann (1913) | Eigenmann (1922) Hoplosternum punctatum Meek & Hildebrand, 1916 X |Maldonado-Ocampo et al. (2006b) Lepthoplosternum ; altamazonicum Reis, 1997 a Gabivet ala 007m) Megalechis picta (Miller & Mojica et al. (2005) | Maldonado- Troschel, 1849) Ocampo et al. (2006a) Megalechis thoracata if (Valenciennes, 1840) Mojica et al. (2005) | Galvis et al. (2007a) Cheyelonas Gondus Gill, 1858) Nijssen and Isbriicker (1983) | Cala (1977) Corydoras agassizii Steindachner, 1876 Castiontge ab) Corydoras ambiacus Cope, 1872 Castro (1987b) Corydoras arcuatus Elwin, 1939 Castro (1987b) Coneanesmeea: Rossel Nijssen and Isbriicker (1983) 1962 Corydoras brevirostris Fraser- ; Brunner, 1947 Galvis et al. (2007a) MES ah irs CZUTIC 11794, 11892 Weitzman, 1961 : ; *Corydoras crypticus Sands, IMCN 5833, IMCN 5834 | CZUT-IC 1995 11837 c Nijssen and Isbriicker (1983) | NRM CLAP CEP OP EIU SED IES eS ase aif |e 26073 [1972222.3251] | NRM 26074 | Isbriicker, 1983 ZMA 119063 Checklist of the freshwater fishes of Colombia: a Darwin Core alternative... 73 Taxa Corydoras elegans Steindachner, 1876 Corydoras esperanzae Castro, 1987 DC Pac Car | References/Collections Castro (1987b) Castro (1987b) | UBJTL MM275 Corydoras evelynae Réssel, 1963 Galvis et al. (2007b) Corydoras fowleri Bohlke, 1950 Corydoras gomezi Castro, 1986 Corydoras habrosus Weitzman, 1960 Mojica (1999). | IMCN 5710 | NRM 16554 Castro (1986) | UBJTL MM536 Castro (1987b) Corydoras leucomelas Eigenmann & Allen, 1942 Castro (1987b) Corydoras loxozonus Nijssen Nijssen and Isbriicker (1983) | ANSP & Isbriicker, 1983 zik 150170 Corydoras melanistius Regan, x Castro (1987b) 1912 Corydoras melanotaenia x |x Regan (1912b) | BMNH 1909.7.23.41 | Regan, 1912 BMNH 1909.7.23.42 Corydoras melini Lonnberg Regan (1912b) | Nijssen and Isbriicker & Rendahl, 1930 (1983) Corydoras metae Eigenmann, x |x far lieexe Eigenmann (1914) | CAS 36447 [ex IU 1914 13451] Corydoras osteocarus Bohlke, x Galvis et al. (20072) 1254 Corydoras pastazensis x Mojica et al. (2005) Weitzman, 1963 ae Corydoras pygmaeus Knaack, x Galvis et al. (2007b) 1966 Corydoras rabauti La Monte, x Castro (1987b) 1941 Corydoras reticulatus Fraser- Brunner, 1938 24 Castro (1987b) Corydoras reynoldsi Myers & x |x x Myers and Weitzman (1960) | SU 52349, Weitzman, 1960 50702 | ZMA 111424 Corydoras semiaquilus Myers and Weitzman (1960) | Arbeldez et Weitzman, 1964 al. (2004) Corydoras septentrionalis 7 Gosline, 1940 Isbriicker (1999) Corydoras simulatus x |x Weitzman and Nijssen 1970) | USNM Weitzman & Nijssen, 1970 197615, 197616, 197667 | ZMA 110384 Corydoras sodalis Nijssen & Isbriicker, 1986 Corydoras splendens Castro (1987b) (Castelnau, 1855) x Mojica et al. (2005) Corydoras trilineatus Cope, BS She Hck oe 1872 Corydoras zygatus Eigenmann - Vere Semon & Allen, 1942 74 Carlos DoNascimiento et al. / ZooKeys 708: 25-138 (2017) . | Mag- ; Taxa DC|EN| ET | Amz| Ori ene Pac| Car| References/Collections Astroblepidae 38 spp (16 added and 3 deleted from 2008) Astroblepus acostai Ardila x |x x Ardila Rodriguez (201 1e) | CAR 473, Rodriguez, 2011 422, | ICN-MHN 17843 Astroblepus ardiladuartei 4 Ardila Rodriaica 015 > nlee.€ xX Ardila Rodriguez (2015a) ee Ardila Rodriguez (2012) | CAR 619, 539, ae vege ee dita ea | x 548, 423, 616 | ICN-MHN 18326 igor IMCN 5260 Astroblepus bellezaensis Ardila ! P Rodriguez, 2015 xX-| X xX Ardila Rodriguez (2015a) Astroblepus cacharas Ardila ‘ : Rodriguez, 2011 xX |X isiS Ardila Rodriguez (2011c) | CAR 460 ee se cl araieuay alle? Io Fowler (1943) | ANSP 70506, 70507-08 ; Eigenmann (1912) | CAS 64658 [ex = 0 cee x | x IU 12708a-c] | FMNH 56071 [ex CM Fiery 4863], 50672 as a as elon - |X] Regan (1912c) | BMNH 1912.3.2.7 Astroblepus curitiensis Ardila ‘ ; Rodriguez, 2015 S a9:¢ x Ardila Rodriguez (2015b) Astroblepus floridablancaensis ; ; Astroblepus frenatus x |x Eigenmann (1918a) | FMNH 58384 [ex Eigenmann, 1918 CM 7380] Astroblepus grixalvii Humboldt, 1805 oe | ie Humboldt (1805) Astroblepus guentheri x |x Boulenger (1887) | BMNH (Boulenger, 1887) 1880.2.26.18-25 paee Si cada arabs dll || Regan (1908) | BMNH 1908.5.29.80 ek Epes orpadi Reset | 4 ling ||| = Regan (1904) | BMNH 1902.5.15.27 Astroblepus itae Ardila x |x x Ardila Rodriguez (2011e) | CAR 555, Rodriguez, 2011 432, 433, 435 | ICN-MHN 7844 Astroblepus jimenezae Ardila x |x x Ardila Rodriguez (2013a) | CAR 470, 542 Rodriguez, 2013 | CIUA 756, 757, 758; re Hepes juriidie Fowles ee Fowler (1944) | ANSP 71431 Eigenmann (1918a) | FMNH 58366 Astroblepus latidens x |x [ex CM 7362], 58367-71, 69816 | CAS Eigenmann, 1918 64659 [ex IU 13678], 64689 [ex IU 13679] ues ESTE ae BOWIE tN Melle Fowler (1919) | ANSP 49368, 49369-84 apeiehis Tearreaias alee * Regan (1904) | BMNH 1899.8.21.10-11 (Regan, 1904) Checklist of the freshwater fishes of Colombia: a Darwin Core alternative... 75 Taxa DC |EN| ET | Amz| Ori ae Pac| Car| References/Collections Astroblepus martinezi Ardila x |x x Ardila Rodriguez (2013a) | CAR 550,421 Rodriguez, 2013 | MBUCV 35674 eae Eigenmann (1918a) | CAS 64692 [ex IU oH tei 2 cr ae Ku NX All 13686] | FMNH 58376 [ex CM 7372], gs > 58377 [ex CM 7373] Astroblepus mojicai Ardila ; ¥ Rodrisues, 2015 XK xX Ardila Rodriguez (2015a) Astroblepus nettoferreirai ’ , Astroblepus nicefori Myers, x |x Pulsars Myers (1932) | SU 24796 | USNM 1932 94178 Astroblepus onzagaensis Ardila ; Rodriguez, 2015 x, | & X Ardila Rodriguez (2015b) Astroblepus orientalis Astroblepus pradai Ardila . ; Rodriguez, 2015 Xt) DE x Ardila Rodriguez (2015b) Astroblepus putumayoensis ; Adda Redsienen 20% xX |X xX Ardila Rodriguez (2015a) Astroblepus rengifoi Dahl, x |x X |Dahl (19600) 1960 Astroblepus retropinnus x |x x Regan (1908) | BMNH 1908.5.29.81-82 (Regan, 1908) | IMCN 295 Eigenmann (1918a) | FMNH 58433 [ex CM 7430], 58372-75, 58427-28, 58431- Astroblepus santanderensis x |x 32 | CAS 47001 [ex IU 13741], 47002 Eigenmann, 1918 [ex IU 13682], 60163 [ex IU 13740], 64696 [ex IU 13683], 64697 [ex IU 13684], 64698 [ex IU 13685] Eigenmann (1912) | AMNH 18667 Astrid leputrafesciat [ex IU 12711] | CAS 64716-17 [ex IU - er tiate ie 8 pes me 12711-12] | FMNH 56076 [ex CM ee eS 4868], 56077-78 | 264718 | USNM 79198 went Eigenmann (1912) | FMNH 56079 [ex at be ee x |X CM 4871], 56080, 69817 | CAS 64718 ark [ex IU 12713] | USNM 79199 Eigenmann (1912) | FMNH 56074 [ex Aga a CM 4866], 56073, 56075 | AMNH ae ae a i 5s eX ale 18666 [ex IU 12710] (1) | CAS 47013 com te [ex CM 4865, IU 12709], 47014 [ex CM 4867a-g, IU 12710a-¢] Astroblepus verai Ardila A ; Rodriguez, 2015 Xa XX x Ardila Rodriguez (2015b) Loricariidae 187 spp (30 added and 24 deleted from 2008) Pseudorinelepis genibarbis alancien nes: LBAO) 4e ||) Ortega-Lara (2016) | Galvis et al. (2007a) Acestridium colombiensis x |x x Retzer (2005) | FMNH 115255, 105169 Retzer, 2005 | INHS 99093 | USNM 381314 76 Carlos DoNascimiento et al. / ZooKeys 708: 25-138 (2017) Taxa *Acestridium dichromum Retzer, Nico & Provenzano, 1999 DC Mag- Ori Cau Pac| Car References/Collections IAvH-P 9946-9948, 10704 Acestridium martini Retzer, Nico & Provenzano, 1999 Hypoptopoma bianale Aquino & Schaefer, 2010 Hypoptopoma brevirostratum Aquino & Schaefer, 2010 Lasso et al. (2005) | IAVH-P 9949-9951 | ICN-MHN 5333, 6328, 12380- 12383,12817,12818, 12384, 12819 Aquino and Schaefer (2010) Aquino and Schaefer (2010) Hypoptopoma gulare Cope, 1878 Hypoptopoma machadoi Aquino & Schaefer, 2010 Hypoptopoma spectabile (Eigenmann, 1914) ~ ~ ~ ~ ~ Aquino and Schaefer (2010) Aquino and Schaefer (2010) Eigenmann (1914) | Aquino and Schaefer (2010) Hypoptopoma steindachneri Boulenger, 1895 Aquino and Schaefer (2010) Hypoptopoma sternoptychum (Schaefer, 1996) Hypoptopoma thoracatum Giinther, 1868 Otocinclus batmani Lehmann A., 2006 Otocinclus huaorani Schaefer, 1997 Otocinclus macrospilus Eigenmann & Allen, 1942 Schaefer (1996) Aquino and Schaefer (2010) Lehmann (2006) | ICN-MHN 6721, 6722 | ANSP 178616 | MCP 28172, 34087 | MHNUC 474 Galvis et al. (2007b) | Maldonado- Ocampo et al. (2013a) Mojica et al. (2005) Otocinclus vestitus Cope, 1872 Mojica et al. (2005) Otocinclus vittatus Regan, 1904 Galvis et al. (2007a) Oxyropsis acutirostra Miranda Ribeiro, 1951 Oxyropsis carinata (Steindachner, 1879) Aquino and Schaefer (2002) Mojica et al. (2005) Oxyropsis wrightiana Eigenmann & Eigenmann, 1889 Parotocinclus eppleyi Schaefer & Provenzano, 1993 Parotocinclus variola Aquino and Schaefer (2002) Maldonado-Ocampo et al. (2006a) Lehmann, Schvambach & Xhl|| oe Lehmann et al. (2015) Reis, 2015 Farlowella acus (Kner, 1854) MPUJ 6067, 6083, 6089, 6095 peter Arbeléez etal. (2004) Retzer and Page (1997) | ANSP 88080 | Farlowella colombiensis Retzer x |x CAS 123733, 169739 [ex CAS 123733] & Page, 1997 ~ ~ ~ ~ ~| x ~ ~ ~ ~ ~ ~ ~ ~ ~ | ICN-MHN 17113 | INHS 32939 [ex ASNP 88080]; Checklist of the freshwater fishes of Colombia: a Darwin Core alternative... fies . | Mag- ‘ Taxa DC |EN| ET | Amz| Ori Ge Pac| Car| References/Collections Farlowella curtirostra Myers, X |Ortega-Lara et al. (2012) 1942 eee gus hesgn, as Regan (1904) | BMNH 1902.5.29.180 Farlowella mariaelenae Martin Salazar, 1964 Retzer and Page (1997) Farlowella mitoupibo Ballen, Urbano-Bonilla & Zamudio,| X | X Ballen et al. (2016b) | MPUJ 8481 2016 Farlowella nattereri Séeiadachnee’ 1910 Xx Retzer and Page (1997) Farlowella oxyrryncha (Kner, a [lee Atbeléez et al. (2004) 1853) id Bogota-Gregory and Maldonado- cea ea x Ocampo (2006a) | ICN-MHN 5748 | IAvH-P 490 *Farlowella taphorni Retzer & Page, 1997 X |IAvH-P 9800, 9801 ek ESA: 4 Retzer and Page (1997) | Cala (1977) Farlowella yarigui Ballen & x |x x Ballen and Mojica (2014) | ICN-MHN Mojica, 2014 17819, 17789, 18889 *Lamontichthys filamentosus (La Monte, 1935) xX IAvH-P 1610, 1617 Lamontichthys lanero x Urbano-Bonilla et al. (2009) Taphorn & Lilyestrom, 1984 pir ena Lamontichthys maracaibero Sturisoma caquetae (Fowler, x |x Fowler (1945) | ANSP 71719 1945) Sturisoma tenuirostre (Steindachner, 1910) ealtt77) Sturisomatichthys aureus x |x Steindachner (1900) | Maldonado-Ocam- (Steindachner, 1900) po et al. (2006b) Sturisomatichthys festivus (Myers, 1942) X |Ortega-Lara et al. (2012) Sturisomatichthys leightoni x |x Regan (1912c) | Eigenmann (1922) | (Regan, 1912) BMNH 1909.7.23.45, 1909.7.23.46-4 Sturisomatichthys panamensis (Eigenmann & Eigenmann, X |Eigenmann (1922) | Regan (1913) 1889) Sturisomatichthys tamanae x |x Regan (1912c) | BMNH 1910.7.11.133, (Regan, 1912) BMNH 1910.7.11.134 CTE GUIICERE GRAD EM | || Seal: Isbricker (1979) | BMNH 1947.7.1.228 Isbriicker, 1979 Crossoloricaria variegata , (Steindachner, 1879) X | Eigenmann (1922) | Regan (1914) Crossoloricaria venezuelae (Schultz, 1944) X | Ortega-Lara et al. (2012) Dasyloricaria filamentosa (Steindachner, 1878) Steindachner (1878) | Londofio-Burbano and Reis (2016) 78 Carlos DoNascimiento et al. / ZooKeys 708: 25-138 (2017) . | Mag- ; Taxa DC|EN|ET | Amz| Ori ene Pac| Car | References/Collections Dasyloricaria latiura _ | x |Figenmann (1912) | FMNH 55115 | (Eigenmann & Vance, 1912) USNM 79219 Dasyloricaria paucisquama Londofio-Burbano & Reis, XC n| OE x Londofio-Burbano and Reis (2016) 2016 Dentectus barbarmatus Martin Salazar, Isbriicker & x Urbano-Bonilla et al. (2009). Nijssen, 1982 Hemiodontichthys - acipenserinus (Kner, 1853) is Moje eteals(2005) Limatulichthys griseus x |x Mojica et al. (2005) | Maldonado-Ocam- (Eigenmann, 1909) po et al. (2006a) she He apse valli c Galvis et al. (2007b) | Galvis et al. (2007a) Linnaeus, 1758 Loricaria nickeriensis “s Isbriicker, 1979 an oneact ay oN) Loggia pide Regan, x Urbano-Bonilla et al. (2009). 1904 Loricariichthys brunneus ; (Hancock, 1828) x Galvis et al. (2007a) *Loricariichthys hauxwelli ee PCIBOE RISERS x Urbano-Bonilla et al. (2009). Rhadinoloricaria laani | ee Nijssen and Isbriicker (1988) | ANSP (Nijssen & Isbriicker, 1988) 131483, 131484, 157750, 157940-41 Rhadinoloricaria listrorhinos x |x ly Nijssen and Isbriicker (1988) | ANSP (Nijssen & Isbriicker, 1988) 131482 Rhadinoloricaria rhami . (Isbriicker & Nijssen, 1983) zi Mey eaeral (2005) Rineloricaria castroi \sbriicker ’ & Nijssen, 1984 X Arbeldez et al. (2004) *Rineloricaria daraha Rapp } Py-Daniel & Fichberg, 2008 X Bogotaé-Gregory et al. (2016) Rineloricaria eigenmanni ; (Pellegrin, 1908) Galvis et al. (2007a) Isbriicker and Nijssen (1979) | FMNH Se eee 83713, 83714-15 | IRSNB 608-609 | Isbriicker & Nissen, 1979 MZUSP [ex IRSNB 608, MZUSP ex sal a ha ale IRSNB 609] | ZMA 114922-23, 115182, 115196-97 | ZSM 25821 Rineloricaria jubata . (Boulenger, 1902) X |Eigenmann (1922) | Regan (1914) Rineloricaria lanceolata ’ (Giinther, 1868) Arbeldez et al. (2004) Rineloricaria magdalenae Sul MG lige Steindachner (1879a) | Eigenmann (Steindachner, 1879) (1922) Rineloricaria rupestris (Schultz, 1944) X | Ortega-Lara et al. (2012) Rineloricaria sneiderni (Fowler, 1944) xX | xX xX Fowler (1944) | ANSP 71433, 71434 Checklist of the freshwater fishes of Colombia: a Darwin Core alternative... Ae) Taxa Spatuloricaria atratoensis DC EN | ET | Amz Ori Mag- Cau ~) 6 Car | References/Collections SP HICTEOHT EA Gaipaatie Xe |X x Fowler (1943) | ANSP 70526, 70527 (Fowler, 1943) Spatuloricaria curvispina (Dahl, 1942) X |X xX Dahl (1942) Spatuloricaria euacanthagenys Isbriicker, 1979 x | 4 a Pomentl ay) Spatuloricaria gymnogaster x |x x x Eigenmann (1912) | Villa-Navarro et al. (Eigenmann & Vance, 1912) (2006) Spatuloricaria fimbriata (eis sinann be Vance 10D) Bs X_ |Eigenmann (1912) | Miles (1947) Spatuloricaria lagoichthys (Schultz, 1944) se) Spatuloricaria phelpsi Schultz, Xf | Ortega:l ara eral (2012) 1944 2, ea Dy Bogota-Gregory and Maldonado-Ocam- ee ee er perc 5) | po (2005) | Cala (1977) | ICN-MHN ar 596 | IAVH-P 2301 Baryancistrus beggini Lujan, ; Arce & Armbruster, 2009 * este 1p eee Baryancistrus demantoides ., : cmt Ajiaco-Martinez et al. (2012c) | Ortega- Werneke, Sabaj Pérez, Lujan X Lara (2016) & Armbruster, 2005 a Chaetostoma aburrense (Posada, 1909) x elliot ~ Posada (1909) Chaetostoma anomalum Regan, 1903 X |Ortega-Lara et al. (2012) Chaetostoma brevilabiatum Dahl, 1942 X |X xX Dahl (1943) Chaetostoma dorsale ; Eisenmann 92> x Urbano-Bonilla et al. (2009). Chaetostoma fischeri ; Steindachner, 1879 AS 2G) S| Bigeninanhy (22) Ardila Rodriguez (2013b) | CAR 633, DEAE IAO ICAI | 58) x 104, 105, 417 | MBUCV 35676 | IMCN Ardila Rodriguez, 2013 5676 Chaetostoma joropo Ballen, Urbano-Bonilla & |e Xx Ballen et al. (2016a) Maldonado-Ocampo, 2016 Chaetostoma lepturum Regan, Regan (1912c) | BMNH 1910.7.11.116- x. ||| xX 1912 118 Chaetostoma leucomelas x |x x lxlx Eigenmann (1918a) | Maldonado-Ocam- Eigenmann, 1918 po et al. (2006b) | Miles (1947) Chaetostoma marginatum Regan, 1904 ~ Regan (1913) 80 Carlos DoNascimiento et al. / ZooKeys 708: 25-138 (2017) Taxa DC|EN|ET | Amz| Ori pd Pac| Car| References/Collections i eo piper asc) | Sa ox X | |Fowler (1944) | ANSP 71432 Chaetostoma palmeri Regan, Regan (1912c) | BMNH 1910.7.11.120- xX | X xX 1912 121 Chaetostoma patiae Fowler, 1945 xX |X xX Fowler (1945) | ANSP 71716, 71717 PAO ITIEE PEERS x |X Regan (1912c) | BMNH 1910.7.11.119 Regan, 1912 Chaetostoma platyrhynchus (Fowler, 1943) Fowler (1943) | ANSP 70512, 70513-15 Chaetostoma setosum x |x Boulenger (1887) | BMNH Boulenger, 1887 1880.2.26.9-10 Chaetostoma sovichthys Chaetostoma tachiraense Schultz, 1944 X |Ortega-Lara et al. (2012) Chaetostoma thomsoni Regan, 1904 X |X xX Regan (1904) | BMNH 1902.5.15.28-30 Chaetostoma vagum Fowler, x |x x Fowler (1943) 1943 Oa SERRE eed Eigenmann (1912) | FMNH 56052 [ex fs Sata: s x i a lik ix CM 4842], 56053-54 | CAS 56745 [ex casei a IU 12698], 74161 [ex IU 12699] Cordylancistrus pijao Provenzano R. & Villa- || OX: X Provenzano R. and Villa-Navarro (2017) Navarro, 2017 Dolichancistrus atratoensis x |x x Dahl (1960c) | ICN-MHN 51, 46 (now (Dahl, 1960) apparently 48) Tiled pale citietic Eigenmann (1916) | FMNH 58350 [ex (Fi hee ; , ‘ Selling x CM 7346], 58351 | CAS 77344 [ex IU oe ae 13661], 77345 [ex IU 13662] Dolichancistrus cobrensis x Ballen and Vari (2012) | ICN-MHN (Schultz, 1944) i 18009 Dolichancistrus fuesslii : (Steindachner, 1911) Calpe 4 x Steindachner (1911) | NUW 48026 ‘Hemiancistrus guahiborum Werneke, Armbruster, Lujan x Maldonado-Ocampo et al. (2006a) & Taphorn, 2005 Hemiancistrus subviridis IMCN 5897, 6094, 6189, 6195, 6196, Werneke, Sabaj, Lujan & X 6284, 6574, 6575 Armbruster, 2005 , : Hypancistrus contradens Armbruster, Lujan & x Maldonado-Ocampo et al. (2008) Taphorn, 2007 Hypancistrus debilittera Armbruster, Lujan & X Ortega-Lara (2016) | IAVH-P 12471 Taphorn, 2007 Checklist of the freshwater fishes of Colombia: a Darwin Core alternative... 81 . | Mag- ’ Taxa DC|EN| ET | Amz| Ori Ge Pac| Car| References/Collections z ee hae aa . Ortega-Lara (2016) | IMCN 5828, 6017, Frasher 2007 6018, 6297, 6298, 6299, 6300 *Hypancistrus inspector Armbruster, 2002 “i Omepaclarsce ie) FOGERTY Maae Maldonado-Ocampo et al. (2008) | Armbruster, Lujan & X IAvEL-P 7044 Taphorn, 2007 i: Leporacanthicus galaxias Isbriicker & Nijssen, 1989 ‘ cecal 4 *Leporacanthicus triactis Isbriicker, Nijssen & Nico, xX IMCN 5772, 5773, 5774 1992 *Leptoancistrus canensis (Meck 8 Hildebrand, 1913) X |Maldonado-Ocampo et al. (2013b) Leptoancistrus cordobensis x |x Dahl and Medem (1964) | IMCN 233, Dahl, 1964 250 Panaque cochliodon me, |X Steindachner (1879b) | NUW 47297-98 (Steindachner, 1879) Panaque nigrolineatus (Peters, 1877) Panaque suttonorum Schultz, 1944 ~ ~ ~ ~ Xx Lujan et al. (2010) | Cala (1977) Lujan et al. (2010) *Panaque titan Lujan, Hidalgo & Stewart 2010 Panagqolus albomaculatus (Kanazawa, 1958) Panagqolus maccus Schaefer & Stewart, 1993 IMCN 6059 Galvis et al. (2007b) Urbano-Bonilla et al. (2009). Peckoltia brevis (La Monte, 1935) Armbruster (2008) *Peckoltia caenosa Armbruster, 2008 IMCN 7061 Peckoltia lineola Armbruster, even 2008 Peckoltia lujani Armbruster, Werneke & Tan, 2015 Armbruster et al. (2015) Peckoltia sabaji Armbruster, 2003 Peckoltia vittata (Steindachner, 1881) Peckoltichthys bachi (Boulenger, 1898) Urbano-Bonilla et al. (2009). Armbruster (2008) Armbruster (2008) Parancistrus Bleeker, 1862 Galvis et al. (2007a) *Pseudoancistrus sidereus Armbruster, 2004 Aphanotorulus ammophilus Armbruster & Page, 1996 Aphanotorulus emarginatus (Valenciennes, 1840) Mf afd) ea hed ~ ~ ~ ~ ~ ~ ~ x |X ~ ~ ~ ~ ~ ~ IMCN 6232, 6234 Maldonado-Ocampo et al. (2008) Bogota-Gregory and Maldonado-Ocam- po (2006a) | Ray and Armbruster (2016) | ICN-MHN 2543 82 Carlos DoNascimiento et al. / ZooKeys 708: 25-138 (2017) . | Mag- ; Taxa DC|EN| ET | Amz| Ori Gi Pac| Car| References/Collections Aphanotorulus horridus em SL" ties Aphanotorulus unicolor (Steindachner, 1908) Ray and Armbruster (2016) Hypostomus annectens : (Regan, 1904) Eigenmann (1922) Hypostomus argus (Fowler, Fowler (1943) | ANSP 70510 1943) Flypostomus carinatus (Steindachner 1881) Maldonado-Ocampo et al. (2008) Hypostomus hemicochliodon Asmbeuster 2003 Lasso et al. (2009) | IAVH-P 8910 ‘Hypostomus holostictus (Regan, 1913) x X | Regan (1913) | BMNH 1913.10.1.57 Hypostomus hondae (Regan, x [Regan (1912c) | Dahl (1955) | 1912) paliereumans C1922) Hypostomus niceforoi (Fowler, x Fowler (1943) | Urbano-Bonilla et al. 1943) (2009). Hypostomus oculeus (Fowler, 1943) Fowler (1943) | ANSP 70518, 70519-20 Hypostomus plecostomoides x Eigenmann (1922) | CAS 82501 [ex IU (Eigenmann, 1922) 15043] | [AVH-P 3045 Hypostomus plecostomus (Linnaeus, 1758) Eigenmann (1922) Hypostomus pyrineusi Mojica et al. (2005) | Maldonado- (Miranda Ribeiro, 1920) Ocampo et al. (2008) *Hypostomus robinii IAvH-P 423, 736, 4957, 5372, 5373, Valenciennes, 1840 5495, 5502, 5504 Hypostomus sculpodon Armbtuster, 2003 sila tabahies Hypostomus varimaculosus es aes Fowler (1945) | ANSP 71707 Hypostomus wilsoni (Risenmani, 1918) x X_ |Eigenmann (1918a) | Dahl (1955) fe, Dele ey SP Fe Steindachner (1878) | MSNG 8856 | ceutacciaeu? |e NMW 42596, 44263-66, 44268), 44294 eindachner, | ZMUC 85 lsorineloricaria villarsi (Liitken, 1874) Ray and Armbruster (2016) Pterygoplichthys gibbiceps (Kner, 1854) Maldonado-Ocampo et al. (2006a) Pterygoplichthys lituratus Hs (Kner, 1854) x Mojica et al. (2005) Pterygoplichthys multiradiatus f (Hancock, 1828) Maldonado-Ocampo et al. (2013a) Pterygoplichthys pardalis . (Castelnau, 1855) Mojica et al. (2005) Pterygoplichthys undecimalis x |x Terr Steindachner (1878) | NMW 47224, (Steindachner, 1878) 47220-22 Checklist of the freshwater fishes of Colombia: a Darwin Core alternative... 83 . | Mag- ; Taxa DC | EN| ET | Amz| Ori Ge Pac| Car| References/Collections : Armbruster and Page (2006) | ICN- Cage x MHN 13455 | FMNH 96959, 101378 | aa eas MSU 2736.2 | USNM 177204 *Prerygoplichthys zuliaensis Weber, 1991 X |Ortega-Lara et al. (2012) Rie PME TAMCATUSLOW CS | Se lige x Fowler (1943) | ANSP 70516 Ancistrus centrolepis Regan, 1913 ORES - | X | X |Regan (1913) | Taphorn et al. (2013) eee HORROS OES, | selllpsy x Fowler (1943) | ANSP 70517 Ancistrus macrophthalmus (Pellegrin, 1912) x Maldonado-Ocampo et al. (2006a) Ancistrus martini Schultz, X |Taphorn et al. (2013) 1944 Ancistrus tolima Taphorn, Armbruster, Villa-Navarro &| X | X Taphorn et al. (2013) | CZUT-IC 4040 Ray, 2013 Ancistrus triradiatus Fideamaim, 1918 be Eigenmann (1918a) Ancistrus vericaucanus Taphorn, Armbruster, Villa- ] X | X Navarro & Ray, 2013 Dekeyseria amazonica Rapp Py-Daniel, 1985 Dekeyseria brachyura (Kner, 1854) Dekeyseria pulchra (Steindachner, 1915) Taphorn et al. (2013) Mojica et al. (2005) xX Galvis et al. (2007a) Dekeyseria scaphirhyncha (Kner, 1854) Maldonado-Ocampo et al. (2006a) Lasiancistrus caucanus Eigenmann, 1912 Lasiancistrus guacharote (Valenciennes, 1840) Lasiancistrus schomburgkii (Giinther, 1864) x x x me ak X Eigenmann (1912) | Fowler (1945) | Dahl (1955) X |Ortega-Lara et al. (2012) Lasiancistrus tentaculatus Armbruster, 2005 Lithoxancistrus orinoco Isbriicker, Nijssen & Cala, 1988 Isbriicker et al. (1988) | ICN-MHN 1200, 1201 | ZMA 119882 | ZMUL 972/3482 *Pseudolithoxus anthrax (Armbruster & Provenzano, 2000) Pseudolithoxus dumus (Armbruster & Provenzano, 2000) Armbruster (2005) Armbruster (2005) | [AvVH-P 5100-5105, 7692, 7982, 7983, 9487, 9595, 9596 IMCN 6601, 6602, 6603,6604, 5874, 5880, 5881, 6063, 6064, 6231 *Pseudolithoxus kelsorum Lujan & Birindelli, 2011 x x x x Xx Maldonado-Ocampo et al. (2006a) IMCN 5876, IMCN 5877 84 Carlos DoNascimiento et al. / ZooKeys 708: 25-138 (2017) Taxa DC|EN|ET | Amz| Ori ao Pac| Car | References/Collections Pseudolithoxus tigris (Arm- bruster & Provenzano, 2000) Cetopsidae 15 spp (4 deleted from 2008) Maldonado-Ocampo et al. (2006a) Cetopsidium morenoi (Fernandez-Yépez, 1972) Cetopsidium pemon Vati, Ferraris & de Pinna, 2005 Cetopsis amphiloxa (Eigenmann, 1914) ~ Vari et al. (2005) Vari et al. (2005) Eigenmann et al. (1914) | Vari et al. * |(2005) Cetopsis baudoensis (Dahl, Eigenmann, 1912 Bunocephalus coracoideus (Cope, 1874) Miles (1945) Mojica et al. (2005) 1960) xX |X - x Dahl (1960c) | ICN-MHN 118, 100 Cetopsis candiru Spix & vy Cetopsis coecutiens : (Lichtenstein, 1819) Vari et al. (2005) | Dahl (1960a) Cetopsis fimbriata Vari, x lx Vari et al. (2005) | USNM 305348, Ferraris & de Pinna 2005 372825, 372826 | ICN-MHN 7272 PORE JIM ae Are ea | Ea |x Fowler (1944) | ANSP 71430 1944) Cetopsis motatanensis (Schultz, 1944) X |Ortega-Lara et al. (2012) Cetopsis orinoco (Schultz, ks 1944) x Cala (1977); Vari et al. (2005) Cetopsis othonops (eecaniantd 912) | x X |Eigenmann (1912) | Vari et al. (2005) Cetopsis umbrosa Vari, ro aedetamsnnos (*{*{ | |x| [| paistat anos **Denticetopsis seducta Vari, Ferraris & de Pinna 2005 x ci aug Helogenes castaneus (Dahl, 1960) Ao es x Dahl (1960a) Helogenes marmoratus x |x Mojica et al. (2005) | Maldonado- Giinther, 1803 Ocampo (2001) Aspredinidae 18 spp (2 added and 2 deleted from 2008) sAnianiiee Pipiiee (isner 4 Friel and Carvalho (2016) 1855) Bunocephalus aleuropsis Cope, 1870 ee | Pee Cardoso (2008) *Bunocephatus aloikae Heder O61 X Cardoso (2008) Bunocephalus colombianus x |x x |lxlx Eigenmann (1912) | Cardoso (2008) | oF fl li x Bunocephalus knerii Steindachner, 1882 Cardoso (2008) Checklist of the freshwater fishes of Colombia: a Darwin Core alternative... 85 Taxa Bunocephalus verrucosus (Walbaum, 1792) Pseudobunocephalus amazonicus (Mees, 1989) DC EN| ET |Amz|O ioae c| Car| References/Collections Bogota-Gregory and Maldonado- Ocampo (2005) Pseudobunocephatus bifidus (Eigenmann, 1942) Pseudobunocephatus lundbergi Friel, 2008 *Pterobunocephalus depressus Hasseman, 1911 Friel (2008) Galvis et al. (2007b) Friel (2008) IAvH-P 674, 8714, 8779, 8814, 8844, 8872, 8903, 11991 Dupouyichthys sapito Schultz, 1944 Mojica et al. (2000) | Ortega-Lara et al. (2012) | Miles (1945) Hoplomyzon papillatus Stewart, 1985 Hoplomyzon sexpapilostoma Taphorn & Marrero, 1990 Xyliphius kryptos Vaphorn & Lilyestrom, 1983 Galvis et al. (2007b) Maldonado-Ocampo et al. (2008) X | Ortega-Lara et al. (2012) Xyliphius lepturus Orcés, 1962 Xyliphius magdalenae Eigenmann, 1912 Xyliphius melanopterus Orcés, 1962 x Cala (1977) | ICN-MHN 1772 Galvis et al. (2007b) | Urbano-Bonilla et al. (2009) Auchenipteridae 46 spp (7 added and 4 deleted from 2008) Centromochlus altae Fowler, 1945 Centromochlus existimatus Mees, 1974 x x Eigenmann (1912) | FMNH 56039 [ex CM 4829} Fowler (1945) | ANSP 71700, 71701-04 | IAVH-P 551 Centromochlus heckelii (De Filippi, 1853) * Centromochlus macracanthus, 2000 Mojica et al. (2005) Lasso et al. (2005) | Mojica et al. (2005) | LAVH-P 966, 1055, 3691, 10964 | ICN-MHN 952, 1927, 2716, 8549, 11950-11952, 12801 TAvH-P 14318 Centromochlus perugiae Steindachner, 1882 Arbeléez et al. (2004) | [AvVH-P 9270-9271, 9284, 9287, 9300-9301, 9314-9315, 9332-9333, 9348-9349, 9361-9362, 9378-9379 Centromochlus reticulatus (Mees, 1974) Mojica et al. (2005) | Galvis et al. (2007a) Centromochlus romani (Mees, 1988) Gelanoglanis stroudi Bohlke, 1980 Urbano-Bonilla et al. (2009). Bohlke (1980) | ANSP 142937, 142938- 39, 142940, 142941 | FMNH 83911, 83912 | MZUSP 14641 Tatia aulopygia (Kner, 1858) IAvH-P 9108, 9134 Tatia dunni (Fowler, 1945) Fowler (1945) | ANSP 71705, 71706 86 Carlos DoNascimiento et al. / ZooKeys 708: 25-138 (2017) Taxa Tatia galaxias Mees, 1974 Tatia gyrina (Eigenmann & Allen, 1942) DC EN] ET | Amz| Ori = ; Pac| Car References/Collections Sarmento-Soares and Martins-Pinheiro (2008) Sarmento-Soares and Martins-Pinheiro (2008) Tatia intermedia (Steindachner, 1877) Sarmento-Soares and Martins-Pinheiro (2008) | IAVH-P 1232 * Tatia marthae Vari & Ferraris, 2013 *Tatia nigra Sarmento-Soares & Martins-Pinheiro, 2008 TAvH-P 12458, 12463, 12473, 12483, 12496, 12861, 12870, 12873 CZUT-IC 4441, 4544, IAVH-P 12862, 12874 * Tatia strigata Soares-Porto, 1995 CZUT-IC 4527, [AvH-P 5634, 10049, 10761, 12577, 12579, 12581-12583 *Ageneiosus dentatus Kner 1857 Ageneiosus inermis (Linnaeus, 1766) Ageneiosus magoi Castillo & Brull, 1989 Galvis et al. (2007a) | Ribeiro et al. (2017) Mojica et al. (2005) | Dahl (1960a) Maldonado-Ocampo et al. (2013a) Ageneiosus pardalis Liitken, 1874 Ageneiosus ucayalensis Castelnau, 1855 Ageneiosus vittatus Steindachner, 1908 Steindachner (1879a) | Dahl (1955) Mojica et al. (2005) ICN-MHN 5476, 5975-5976, 5978, 6605, 9178 *Asterophysus batrachus Kner, 1858 Tympanopleura atronasus (Eigenmann & Eigenmann, 1888) x x x Xx x x Ortega-Lara (2016) | IMCN 5528, 5529, 5530, 5964, 6116, 6117, 6118, 6119, 6120, 6121, 6122, 6123, 6124, 6433, 6434 Mojica et al. (2005) Tympanopleura brevis (Steindachner, 1881) Tympanopleura piperata Eigenmann, 1912 *Auchenipterichthys coracoideus Eigenmann & Allen, 1942 Auchenipterichthys longimanus (Ginther, 1864) x Xx x Mojica et al. (2005) Mojica et al. (2005) Galvis et al. (2007b) Bogotd-Gregory and Maldonado-Ocam- po (2006a) | Maldonado-Ocampo et al. (2006a) | IAVH-P 255, 5260 Auchenipterichthys punctatus (Valenciennes, 1840) Auchenipterus ambyiacus Fowler, 1915 Auchenipterus nuchalis (Spix & Agassiz, 1829) Bogota-Gregory and Maldonado- Ocampo (2006a) | [AVH-P 254, 2393, 10008-10019 Ferraris and Vari (1999) Maldonado-Ocampo et al. (2008) | Mojica et al. (2005) Entomocorus gameroi Mago- Leccia, 1984 Se _|L__1L_ Lal ELI Las ee pee x Reis and Borges (2006) Checklist of the freshwater fishes of Colombia: a Darwin Core alternative... 87 . | Mag- ; Taxa DC|EN|ET | Amz| Ori Ge Pac| Car| References/Collections Epapterus dispilurus Cope, x Mojica et al. (2005) 1878 Liosomadoras morrowi x Bogota-Gregory and Maldonado-Ocam- Fowler, 1940 po (2005) Liosomadoras oncinus ; (Jardine, 1841) x Galvis et al. (2007b) Pseudepapterus cucuhyensis : Bohlke, 1951 xX Bohlke (1951) Pseudepapterus hasemani ss (Steindachner, 1915) Mojica et al. (2005) Tetranematichthys wallacei Vari and Ferraris (2006) | [AvH-P 9084 Vari & Ferraris, 2006 tl gue ¥ Trachelyichthys decaradiatus Mees, 1974 Galvis et al. (2007a) Trachelyopterichthys anduzei Ferraris & Fernandez, 1987 pialdonade: Oanpecvar nz Qed) Trachelyopterichthys taeniatus Bogota-Gregory and Maldonado-Ocampo (Kner, 1858) (2006a) | Cala (1977) | CAS-SU 42795 Eigenmann (1916) | FMNH 57695 [ex Trachelyopterus fisheri x |x CM 6667a], 57696-98 | CAS 52136 [ex (Eigenmann, 1916) IU 13496], 57937 [ex IU 13495], 57938 [ex IU 13497] | USNM 76929 Trachelyopterus galeatus u (Linnaeus, 1766) Mojica et al. (2005) | Galvis et al. (2007a) Trachelyopterus insignis x |x Steindachner (1878) | Mojica et al. (Steindachner, 1878) Trachelyopterus peloichthys (Schultz, 1944) Trachycorystes trachycorystes (Valenciennes, 1840) (2006a) IAvH-P 12863, 12876, 12889 Doradidae 56 spp (7 added and 4 deleted from 2008) Acanthodoras cataphractus (Linnaeus, 1758) Bogota-Gregory and Maldonado- Ocampo (2006a) | Galvis et al. (20072) | ICN-MHN 360, 2772, 4183, 5402 *Acanthodoras depressus (Steindachner, 1881) Acanthodoras spinosissimus (Eigenmann & Eigenmann, 1888) CZUT-IC 4312 Mojica et al. (2005) | Maldonado-Ocam- po et al. (2006a) Agamyxis albomaculatus (Peters, 1877) Galvis et al. (2007a) Agamyxis pectinifrons (Cope, 1870) *Astrodoras Bleeker, 1862 Amblydoras affinis (Kner, 1855) Amblydoras bolivarensis (Fernandez-Yépez, 1968) Mojica et al. (2005) Roa-Fuentes et al. (2010) Mojica et al. (2005) | [AvVH-P 747, 1075, 1181, 2807 Maldonado-Ocampo et al. (2006a) (Humboldt, 1821) Centrodoras brachiatus (Cope, 1872) * Centrodoras hasemani (Steindachner, 1915) Doras phlyzakion Sabaj Pérez Bogota-Gregory and Maldonado-Ocam- po (2006a) | IAvH-P 405, 479 CZUT-IC 14832 88 Carlos DoNascimiento et al. / ZooKeys 708: 25-138 (2017) Taxa DC | EN| ET | Amz| Ori at Pac| Car| References/Collections Amblydoras gonzalezi Amblydoras monitor (Cope, * 1872) xX Mojica et al. (2005) Amblydoras nauticus (Cope, - 1874) x Mojica et al. (2005) Anadoras grypus (Cope, 1872) x Mojica et al. (2005) Anadoras regani x Bogota-Gregory and Maldonado-Ocampo (Steindachner, 1908) (2005) Hypodoras forficulatus x Figenmiannsit925 x Mojica et al. (2005) Physopyxis ananas Sousa & Rapp Py-Daniel, 2005 Beall ee Maldonado-Ocampo et al. (2008) Physopyxis lyra Cope, 1871 X Mojica et al. (2005) Pee Deck Es SC) Correa (2003) | Galvis et al. (2007a) Anduzedoras oxyrhynchus enna pe) |e swe RR ae ne los Xx Humboldt and Valenciennes (1821) x x & Brindelli, 2008 Sabaj Pérez and Birindelli (2008) Doraops zuloagai Schultz, 1944 x Ortega-Lara et al. (2012) Hassar orestis (Steindachner, 1875) Hemidoras stenopeltis (Kner, 1855) Birindelli et al. (2011) | IAVH-P 6046 Mojica et al. (2005) Hemidoras stuebelii (Steindachner, 1882) Mojica et al. (2005) Leptodoras acipenserinus (Giinther, 1868) Leptodoras juruensis Boulenger, 1898 Leptodoras linnelli Eigenmann, 1912 Maldonado-Ocampo et al. (2008) Mojica et al. (2005) TAvH-P 15132 Leptodoras nelsoni Sabaj Pérez, 2005 *Leptodoras rogersae Sabaj Pérez, 2005 *Lithotodoras dorsalis (Valenciennes, 1840) Sabaj (2005) | IAVH-P 904 CZUT-IC 9529 CZUT-IC 14500 Megalodoras uranoscopus (Eigenmann & Eigenmann, 1888) Li Lie _ ee E eee ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~< ~ Mojica et al. (2005) Checklist of the freshwater fishes of Colombia: a Darwin Core alternative... 89 Taxa Nemadoras cristinae Sabaj Pérez, Arce H., Sousa & Birindelli, 2014 DC EN | ET | Amz Ori Mag- Cau Pac Car | References/Collections Sabaj Pérez et al. (2014) Nemadoras elongatus (Boulenger, 1898) Sabaj Pérez et al. (2014) Nemadoras hemipeltis (Eigenmann, 1925) Nemadoras humeralis (Kner, 1855) Orinocodoras eigenmanni Myers, 1927 Sabaj Pérez et al. (2014) Sabaj Pérez et al. (2014) Maldonado-Ocampo (2001) Opsodoras boulengeri Steindachner, 1915 Opsodoras morrisi Eigenmann, 1925 Ossancora punctata (Kner, 1853) Mojica et al. (2005) Bogota-Gregory and Maldonado-Ocam- po (2006a) Birindelli and Sabaj Pérez (2011) Oxydoras niger (Valenciennes, 1821) Mojica et al. (2005) | Maldonado-Ocam- po et al. (2006a) Oxydoras sifontesi Fernandez- Yépez, 1968 Maldonado-Ocampo et al. (2006a) | CZUT-IC 9380 Platydoras armatulus ae (Valenciennes, 1840) Profstaver al (2003) Platydoras hancockii (Valenciennes, 1840) Cala (1977) Pterodoras granulosus (Valenciennes, 1821) Pterodoras rivasi (Fernandez- Yépez, 1950) *Rhinodoras boehlkei Glodek, Whitmire & Orcés V., 1976 Mojica et al. (2005) Birindelli (2014) CZUTAIG 122725.12318 Rhinodoras gallagheri Sabaj, Taphorn & Castillo G., 2008 Rhinodoras thomersoni Taphorn & Lilyestrom, 1984 Maldonado-Ocampo et al. (2008) X |Ortega-Lara et al. (2012) Tenellus leporhinus (Eigenmann, 1912) Sabaj Pérez et al. (2014) Tenellus ternetzi (Eigenmann, 1925) Tenellus trimaculatus (Boulenger, 1898) Trachydoras microstomus (Eigenmann, 1912) Sabaj Pérez et al. (2014) TAvH-P 10752 Trachydoras nattereri (Steindachner, 1881) Trachydoras steindachneri (Perugia, 1897) Mojica et al. (2005) 90 Carlos DoNascimiento et al. / ZooKeys 708: 25-138 (2017) . | Mag- ; Taxa DC | EN| ET | Amz| Ori Gi Pac| Car | References/Collections Ariidae 1 sp Notarius bonillai (Miles, Miles (1945) | Acero and Betancur-R 1945) | alee . * | 2006) Heptapteridae 52 spp (14 added and 7 deleted from 2008) *Brachyrhamdia imitator *Brachyrhamdia meesi Sands &¢ Black, 1985 xX TAvH-P 11146 *Brachyrhamdia thayeria Slobodian & Bockmann, x Galvis et al. (2007b) 2013 ; Eigenmann (1922) | CAS 63607 [ex IU iced Sea ha x |X x 15004] | FMNH 55212 [ex CM 3923], =i a 55213 [ex CM 3924] Cetopsorhamdia molinae Miles, 1943 Sale. 2 Miles (1943) Cetopsorhamdia nasus x |x x Eigenmann (1916) | FMNH 58126 [ex Eigenmann & Fisher, 1916 CM 7124] Cetopsorhamdia orinoco Schultz, 1944 xX Urbano-Bonilla et al. (2009). Cetopsorhamdia picklei Schultz, 1944 bs X |Ortega-Lara et al. (2012) Ch Eigenmann (1922) | FMNH 55140 [ex e Hed: oa BE |x Ee CM 3841], 55141 [?CM 3842] | CAS Sore a 75751 [ex IU 15019] Gladioglanis conquistador Lundberg, Bornbusch & x Galvis et al. (2007b) Mago-Leccia, 1991 Seen Mes Maldonado-Ocampo et al. (2006a) | Aen ie se er 88 CZUT-IC 5091, 5126 1989 Goeldiella eques (Miiller & Correa (2003) | Maldonado-Ocampo Troschel, 1848) (2001) Heptapterus panamensis Bussing, 1970 X | AvH-P 7273-7274 Imparfinis microps x |x Eigenmann (1916) | FMNH 57793 Eigenmann & Fisher, 1916 [ex CM 6776 not 8778] Imparfinis nemacheir . (Hist tmani 8 Eehees1916) X |Eigenmann (1916) | Eigenmann (1922) Imparfinis pristos Mees & Mees and Cala (1989) | ICN-MHN Cala, 1989 1401, 1402 | RMNH 30544 Imparfinis pseudonemacheir Mees & Cala, 1989 Xx Mees and Cala (1989) ee sprpaete Beeaiy || Se | X | |Regan (1913) | BMNH 1913.10.1.41 Imparfinis stictonotus (Fowler, 1940) xX Fowler (1945) Checklist of the freshwater fishes of Colombia: a Darwin Core alternative... 91 . | Mag- ' Taxa DC |EN| ET | Amz| Ori bt Pac| Car| References/Collections Imparfinis timana Ortega- Lara, Milani, DoNascimiento, Valli ete isid tes x |X Ortega-Lara et al. (2011) | [AVH-P 10696 Ocampo, 2011 Imparfinis usmai Ortega-Lara, plant Dalasi: a IX Ortega-Lara et al. (2011) | IMCN 4812 Villa-Navarro & Maldonado- aa Ocampo, 2011 Mastiglanis asopos Beale tannn 1994 Galvis et al. (2007b) Myoglanis koepckei Chang, Galvis et al. (2007b) 1990 NeRuRIgATHS TA4n4? |x x Schultz (1944a) | USNM 121251 (Schultz, 1944) *Nemuroglanis pauciradiatus Ferraris, 1988 IAvH-P Pariolius armillatus (Mees, 1987) *Phenacorhamdia anisura (Mees, 1987) Phenacorhamdia macarenensis IAvH-P 8680, 8935, 9003, 9087, 9113, 9389, 9433 TAvH-P 12894 Dahl, 1961 ee |e x Dahl (1961) Phenacorhamdia nigrolineata Zarske, 1998 x Maldonado-Ocampo et al. (2008) *Phenacorhamdia provenzanoi DoNascimiento CZUT-IC 6351, 6881, 6988 & Milani, 2008 *Phenacorhamdia taphorni DoNascimiento & Milani, TAvH-P 7932, 9254, 9594, 10727 2008 Pimelodella chagresi (Steindachner, 1876) Fowler (1944) | Eigenmann (1922) Pimelodella conquetaensis Ahl, 1925 x | X Ahl (1925) | ZMB 32030 Pimelodella cristata (Miiller Arbelaez et al. (2004) | Urbano-Bonilla et & Troschel, 1848) al. (2009). *Pimelodella cruxenti Fernandez-Yépez, 1950 Soe eas i se a PUGET ER 5 | X [Regan (1913) | BMNH 1913.10.1.37-40 Pimelodella figueroai Dahl, x |x Dahl (1961) 1961 Pimelodella floridablancaensis x |x Ardila Rodriguez (2017a) Ardila Rodriguez, 2017 Pimelodella geryi Hoedeman, 1961 Pimelodella gracilis (Valenciennes, 1835) Arbeldez et al. (2004) Bogota-Gregory and Maldonado-Ocam- 1847, 1967, 3724, 3725-3729, 10004- 10005 | ICN-MHN 1816, 3731, 1909, 2179, 5512, 5654 Zuanon, Rapp Py-Daniel & Jégu, 1993 Brachyplatystoma Mojica et al. (2005) filamentosum (Lichtenstein, I 4)| OK Ajiaco-Martinez et al. 2012a) 1819) a UL led allie € Ramirez-Gil et al. (2012a) | Cala (1977) (Boulenger, 1898) Brachyplatystoma platynemum Boulenger, 1898 Brachyplatystoma rousseauxii ~ ~ Ramirez-Gil et al. (2012b) (Castelnau, 1855) ee a, Agudelo-Cérdoba et al. (2012) Brachyplatystoma tigrinum ; (Britski, 1981) x Mojica et al. (2005) Brachyplatystoma vaillantii ln Asso! Nunn oot MAD (Valenciennes, 1840) Platynematichthys notatus (Jardine, 1841) Mojica et al. (2005) | Maldonado-Ocam- 93 Carlos DoNascimiento et al. / ZooKeys 708: 25-138 (2017) Taxa DC|EN|ET | Amz| Ori ae Pac| Car| References/Collections Pimelodella grisea (Regan, Pimelodella linami Schultz, x Galvis et al. (2007a) 1944 Pimelodella macrocephala xlxlx x Miles (1943) | MCZ 35876 | USNM (Miles, 1943) 120157 Piidlodilie wet Eigenmann (1917b) | CAS [ex IU 13768] as agit ee x | FMNH 58441 [ex CM 7441 or 7141], owes 58442 | ICN-MHN 17128 Pimelodella modestus ; (Giinther, 1860) x Eigenmann (1922) Pimelodella odynea Schultz, Sy | Sikepact ara’erall (0012) 1944 renames xix] | fx] | fotos Pimelodella reyesi Dahl, 1964 | X | X a a ee Dahl and Medem (1964) *Rhamdia guatemalensis , (Giinther, 1864) X | X | X | Hernandez et al. (2015) Rhamdia laukidi Bleeker, x HeratndesietalcOOis) 1858 Bogota-Gregory and Maldonado- Rhamdia quelen (Quoy & Ocampo (2006a) | Eigenmann (1922) | Gaymard, 1824) IAvH-P 6182, 6223 | ICN-MHN 5029, 6579, 6591 Rhamdia saijaensis Rendahl, x |x Rendahl (1941) 1941 Pimelodidae 53 spp (3 added and 4 deleted from 2008) Aguarunichthys inpai xX xX xX xX xX Calophysus macropterus (Lichtenstein, 1819) ~ ~ ~ ~ po et al. (2004) Dahl (1961) | Mojica et al. (2005) | IAvH-P 2210 | ICN-MHN 7889, 8158 Checklist of the freshwater fishes of Colombia: a Darwin Core alternative... 93 Taxa DC|EN|ET|Amz|O ae "| Pac| Car] References/Collections Cheirocerus abuelo (Schultz, 1944) le X | Ortega-Lara et al. (2012) Cheirocerus goeldii (Steindachner, 1908) Mojica-er al. (2005) Duopalatinus peruanus Eigenmann & Allen, 1942 = Mae Teh ne oe 4 CZUT-IC 9407, 9420, 9553, 9660, aia 9708, 9727, 9733, 9745, 9842 Nass, 1991 Hemisorubim platyrhynchos x Maldonado-Ocampo et al. (2006a) | (Valenciennes, 1840) Mojica et al. (2005) Fe ak aaa X Mojica et al. (2005) | Galvis et al. (2007a) Spix & Agassiz, 1829 J , : Hypophthalmus fimbriatus ¥ Kner, 1858 Mojica et al. (2005) Hypophthalmus marginatus Mojica et al. (2005) | Lasso et al. (2005) | Valenciennes, 1840 ICN-MHN 8228 Hypophthalmus oremaculatus Mojica et al. (2005) | Littmann et al. Nani & Fuster, 1947 (2015) on aren ate aU Mojica et al. (2005) | Eigenmann (1922) Leiarius perruno (Schultz, 1944) Ortega-Lara et al. (2012) Leiarius pictus (Miiller & Lasso et al. (2005) | Correa (2003) | Troschel, 1849) IMCN 842 Megalonema orixanthum Lundberg and Dahdul (2008) | ANSP Lunberg & Dahdul, 2008 187449 (ex ANSP 148143) Megalonema platycephalum lx Dahl (1961)} Eigenmann, 1912 Megalonema psammium Schultz, 1944 X |Ortega-Lara et al. (2012) Eigenmann (1912) | FMNH 56032 [ex CM 4822] | AMNH 5340 | BMNH Mea th 1920.12.20.112-113, 1924.3.3.83-84 | ie tigi ie ; byt ane ale x CAS 63674-75 [ex IU 12681-82] | FMNH ee 10251, 10284-89, 77909, 56032-33, 69818, 95984 | MCZ 30961 | UMMZ 190406 | USNM 76930, 79222, 167852 Phractocephalus hemioliopterus (Bloch & Mojica et al. (2005) | Dahl (1961) Schneider, 1801) Pimelodina flavipinnis Lasso et al. (2005) | Mojica et al. (2005) | Steindachner, 1877 ICN-MHN 6874, 8143 Lasso et al. (2005) | IAVH-P 256, 503, Pimelodus albofasciatus Mees, 836, 477, 2150, 2677, 2731, 6023, 8754, 1974 8821, 8850, 8913 | ICN-MHN 1259, 1951, 1798, 3725, 3726 | 3822, 3386 - Mojica et al. (2005) | Galvis et al. (2007a) haere 40 | Mojica et al. (2006a) | Maldonado- Reg Ocampo et al. (2013b) 94 Carlos DoNascimiento et al. / ZooKeys 708: 25-138 (2017) Taxa DC |EN| ET | Amz| Ori ae Pac| Car| References/Collections Pimelodus coprophagus Schultz, 1944 Pimelodus crypticus Villa- Navarro & Cala, 2017 BS Villa-Navarro et al. (2017) Pimelodus garciabarrigai x Dahl (1961) | ICN-MHN 744, Dahl, 1960 ICN-MHN 410 Pimelodus grosskopfii x Steindachner (1879b) | Steindachner, 1879 NMW 45781-45782 ip (iia aa or X_ |Mojica (1999) | ICN-MHN 2150 . Bsa ECD LED Mojica et al. (2005) | Dahl (1961) Pimelodus pictus r Steindachner, 1877 Mojica et al. (2005) | Galvis et al. (2007a) Pimelodus punctatus (Meek x Maldonado-Ocampo et al. (2013b) | & Hildebrand, 1913) : Villa-Navarro et al. (2017) Pimelodus yuma Villa- : Navarro & Acero P, 2017 x |X X X | Villa-Navarro et al. (2017) Pinirampus pirinampu (Spix Mojica et al. (2005) | Maldonado- & Agassiz, 1829) Ocampo et al. (2006a) Platysilurus malarmo Schultz, x X |Ortega-Lara et al. (2012) 1944 ee toad Valiant Mojica et al. (2005) | Galvis et al. (2007a) Platystomatichthys sturio ‘ (Kner, 1858) Mojica et al. (2005) Pseudoplatystoma ; , s dialennhire Butane: x Buitrago-Sudrez and Burr (2007) | Suarez & Burr, 2007 Pseudoplatystoma metaense Buitrago-Suarez & Burr, 2007 CAS 19165 | FMNH 56278, 59234 Buitrago-Sudrez and Burr (2007) | Pseudoplatystoma orinocoense Buitrago-Suarez & Burr, 2007 Pseudoplatystoma punctifer (Castelnau, 1855) - a ANSP 146858, 128135, 149541 xX x Buitrago-Sudrez and Burr (2007) Buitrago-Sudrez and Burr (2007) Pseudoplatystoma tigrinum (Valenciennes, 1840) Buitrago-Sudrez and Burr (2007) Sorubim cuspicaudus Littmann, Burr & Nass, 2000 Littmann et al. (2000) | AUM 28756 | EMNH 56223, 60305, 107492 | IAvH-P 11809 | INHS 35428; Sorubim elongatus Littmann, Burr, Schmidt & Isern, 2001 Littmann et al. (2001) Sorubim lima (Bloch & Schneider, 1801) Littmann (2007) | Dahl (1961) Sorubim maniradii Littmann, Burr & Buitrago- x Mojica et al. (2005) Suarez, 2001 Sorubimichthys planiceps x|xlx Mojica et al. (2005) | Dahl (1961) | ICN- (Spix & Agassiz, 1829) MHN 7880, 9938, 17351 acanthochiroides (Giinther, Checklist of the freshwater fishes of Colombia: a Darwin Core alternative... 95 . | Mag- , Taxa DC |EN| ET | Amz| Ori ee Pac| Car| References/Collections Zungaro Zungaro nie ls || na Pseudopimelodidae 12 spp (1 added from 2008) Batrochoglanis - | + |Ortega-Lara et al. (2012) 1942) Batrochoglanis raninus (Valenciennes, 1840) Xx Bogota-Gregory and Maldonado- Ocampo (2005) Batrochoglanis transmontanus (Regan, 1913) Batrochoglanis villosus (Eigenmann, 1912) Regan (1913) | Maldonado-Ocampo et al. (2013b) Lasso et al. 2004, Bogotd-Gregory and Maldonado-Ocampo 2006, Maldonado- Ocampo et al. 2008, Lasso et al. 2009 | [AVH-P 5705, 10059, 10060, 10061, 10062, 10063, 10064, 10065, 10066, 10743, 10798, 12466, 12467, 12481, 12867, 12880, 12913 Cephalosilurus apurensis (Mees, 1978) Lasso et al. (2005) | IAVH-P 3731 Cruciglanis pacifici Ortega- Ortega-Lara and Lehmann (2006) | Lara & Lehmann, 2006 x IMCN 2359 | IAvH-P 7505 Microglanis iheringi Gomes, 1946 | Cala (1977) Microglanis poecilus Mojica et al. (2005) | Maldonado- Eigenmann, 1912 Ocampo et al. (2006a) pens pega Mee Lasso et al. (2005) | ICN-MHN 1912 Lasso et al. (2005) | Ortega-Lara et al. Re inaeal e ee X | (2012) | IAVH-P 3371, 9198-9199, aes 11294, 12291 | ICN-MHN 772, 6582 Pseudopimelodus schultzi i (Dahl, 1955) xX. | XOX X | Dahl (1955) | Mojica et al. (2006a) Rhyacoglanis annulatus . , Shibatta & Vari, 2017 Shibatta and Vari (2017) Batrachoidiformes 3 spp Batrachoididae jes nf | ioe eS Bengals | Seales X |Rendahl (1941) | NRM 10651 Daector quadrizonatus : X |X - | X |Eigenmann (1922) | FMNH [ex CM 3921] (Eigenmann, 1922) Thalassophryne amazonica Steindachner, 1876 Gobiiformes 1 sp (1 deleted from 2008) Mojica et al. (2005) Eleotridae Microphilypnus ternetzi Myers, 1927 Caires and Figueiredo (2011) | Galvis et al. (2007a) 96 Carlos DoNascimiento et al. / ZooKeys 708: 25-138 (2017) T . | Mag- : axa DC|EN|ET | Amz| Ori Cau Pac| Car| References/Collections Synbranchiformes 1 sp Synbranchidae SSA Mojica et al. (2005) | Fowler (1943) | Gal- Synbranchus marmoratus vis et al. (2007a) | Steindachner (1880) Bloch, 1795 | Maldonado-Ocampo et al. (2013b) | Regan (1913) Pleuronectiformes 5 spp (2 added and 2 deleted from 2008) Achiridae Achirus novoae Cervigén, x Mald 1982 onado-Ocampo et al. (2006a) *Apionichthys nattereri . ees x] ff | Mei cat coos a . Ramos (2003b) | ANSP 176081, 163851- phe ee anos X 53,163854, 163855-56 | MZUSP 52055 | UEPB 3596 Hepoih: ik Mojica et al. (2005) | Lasso et al. (2009) Dea tei, ay oe x |x | IMCN 2657, 2662 | IAVH-P 10784- (Giinther, 1862) 10787 ee | (Supe Duplain etal, (2012) | BMNH Munroe, 2012 Ovalentaria incertae sedis Polycentridae 1 sp Monocirrhus polyacanthus Heckel, 1840 #915, FO M19, Arbeléez et al. (2004) | Cala (1977) Cichliformes 94 spp (8 added and 30 deleted from 2008) Cichlidae Cichla intermedia Machado- Allison, 1971 Cichla monoculus Spix & Agassiz, 1831 ~ ~< ~ ~ Maldonado-Ocampo (2001) Mojica et al. (2005) | Maldonado-Ocam- po et al. (2006a) Cichla orinocensis Humboldt, 1821 Cichla temensis Humboldt, 1821 Chaetobranchus flavescens Heckel, 1840 Kullander and Ferreira (2006) Galvis et al. (2007b) | Cala (1977) Kullander (1986) | Galvis et al. (2007a) Acarichthys heckelii (Miller & Troschel, 1849) Kullander (1986) Biotoecus dicentrarchus Kullander, 1989 Crenicara punctulatum (Ginther, 1863) Kullander (1989a) | ICN-MHN 1400 | EMNH 85650, 105089 | NRM 16662, 37078 Mojica et al. (2005) Checklist of the freshwater fishes of Colombia: a Darwin Core alternative... 97 Taxa DC |EN| ET | Amz| Ori ae Pac| Car | References/Collections ae Bogota-Gregory and Maldonado- ee aa Eapensnant Ocampo (2005) | Maldonado-Ocampo et al. (2006a) sha PEDERI COPE Correa (2003) | Galvis et al. (2007a) Crenicichla geayi Pellegrin, 1903 Regan (1905) Crenicichla johanna Heckel, Mojica et al. (2005) 1840 Crenicichla lenticulata Heckel, 1840 Correa (2003) | Cala (1977) A Fowler (1943) | LAVH-P 389, 2327, 4893, sl Metres ede, 10110 | ICN-MHN 867, 877, 884, 1553, 1668, 2286 Crenicichla monicae Kullander & Varela, 2015 Kullander and Varella (2015) Crenicichla reticulata (Heckel, 1840) Kullander (1986) Crenicichla strigata Giinther, Bogota-Gregory and Maldonado-Ocam- 1862 po (2006a) | ICN-MHN 2594, 10233 -_ . Ploeg (1991) | RMNH 31622, 31623 | Crenicichla sveni Ploeg, 1991 vel lst 4 eal wl 7MA, 120388 *Crenicichla zebrina Montana, Lépez-Fernandez X IMCN 6476 & Taphorn 2008 Dicrossus filamentosus (Ladiges, 1958) | Cala (1977) Dicrossus gladicauda Schindler and Staeck (2008) | MTDF Schindler & Staeck, 2008 31312, 33313-18 Apistogramma agassizii (Steindachner, 1875) Rallonden(1286) ‘sey ek Kullander (2004) | UF 33670, 128785, i a ie x | x 33687, 33717 | CAS 50638 | ICNM- cae NHN 6735 | NRM 27040, 33375, 36102 Apistogramma bitaeniata Pellegrin, 1936 xX Kullander (1986) Apistogramma cacatuoides Eedecane lel a6 Kullander (1986) Apistogramma cruzi Kullander, 1986 | nce Apistogramma diplotaenia Bogoté-Gregory and Maldonado-Ocampo Kullander, 1987 (2005) Apistogramma eunotus Kullander, 1981 Kullander (1986) | NRM 17784 *Apistogramma flabellicauda CZUT-IC 1494, IAVH-P 10087, 10089- Mesa S. & Lasso, 2011 10092, 10094-10098 Apistogramma hoignei : Meinken, 1965 Urbano-Bonilla et al. (2009). Apistogramma hongsloi rah eae Bs cx Kullander (1979) | NRM 11234, 11235-39 98 Carlos DoNascimiento et al. / ZooKeys 708: 25-138 (2017) . | Mag- ; Taxa DC |EN| ET | Amz| Ori ene Pac| Car| References/Collections Apistogramma iniridae x |x x |x Kullander (1979) | Bogota-Gregory and Kullander, 1979 Maldonado-Ocampo (2005) Apistogramma lineata Mesa S. & Lasso, 2011 x |X X Mesa S and Lasso (2011b) OE eI Kullander (1979) | AMNH 22733 | eat i eae Te Vee |e x FMNH 55218 [ex CM 3930] | ICN- ane MHN 17117 | NRM 11240, 11241-44 Apistogramma megaptera Mesa & Lasso, 2011 X Mesa Salazar and Lasso (201 1a) Apistogramma minima Mesa x Mesa S and Lasso (2011b) | LAvH-P S. & Lasso, 2011 11876 Apistogramma piaroa Mesa S. x Mesa S and Lasso (201 1b) & Lasso, 2011 Apistogramma velifera Staeck, x Mesa $ and Lasso (201 1b) 2003 Be a igi Xx Kullander (1979) | NRM 11231, 11232 Kullander, 1979 on ; Apistogrammoides pucallpaensis Meinken, 1965 * Sal a i Biotodoma cupido (Heckel, xt 1840) x Mojica et al. (2005) Biotodoma wavrini (Gosse, x |x Maldonado-Ocampo et al. (2006a) | 1963) Arbeldez et al. (2004) Geophagus abalios Lépez- x Bogota-Gregory and Maldonado-Ocam- Fernandez & Taphorn, 2004 po (2005) | Galvis et al. (2007a) Geophagus crassilabris Steindachner, 1876 x HON SiN Geophagus dicrozoster Lépez- Fernndez & Taphorn, 2004 x Maldonado-Ocampo et al. (2006a) Geophagus megasema Heckel, x Bogota-Gregory and Maldonado-Ocam- 1840 po (2005) aoa ae ere | || ete X | X [Regan (1912a) | Eigenmann (1922) Geophagus steindachneri ; f ecnianeehorhicebeind: x x Eigenmann (1910) | Ortega-Lara et al. (2012) 1910 Geophagus surinamensis x Correa (2003) (Bloch, 1791) ad Geophagus taeniopareius Kullander & Royero, 1992 x Maldonado-Ocampo et al. (2006a) CGasbhapastenemilae Mipee Bogota-Gregory and Maldonado-Ocam- Fernandez & Taphorn, 2004 pei200>) | Malignady Ocampornd ; Bogota-Gregory (2007) Mikrogeophagus ramirezi (Myers & Harry, 1948) a7) Satanoperca daemon (Heckel, Correa (2003) | Maldonado-Ocampo et 1840) al. (2006a) aes jurupari (Heckel, x Kullander (1986) Checklist of the freshwater fishes of Colombia: a Darwin Core alternative... 99 Taxa DC|EN|ET | Amz| Ori ad Pac| Car | References/Collections Satanoperca mapiritensis : (Fernandez-Yépez, 1950) Galvisireal(20073) nae ocellatas SSASstt, Mojica et al. (2005) | Galvis et al. (2007a) Acaronia nassa (Heckel, Bogota-Gregory and Maldonado-Ocam- 1840) po (2006a) | [AvH-P 5308 Acaronia vultuosa Kullander, Kullander (1989b) 1989 Aequidens diadema (Heckel, Arbeldez et al. (2004) | Maldonado- 1840) Ocampo et al. (2006a) aa ince ets Eigenmann (1922) | CAS 66884 [ex IU i a Se 13967], 66983 [ex IU 13967] | I[AvH-P 1922 3162 Aequidens tetramerus 4 (Heckel, 1840) | Mojica et al. (2005) Andinoacara biseriatus yu | 36 X |Regan (1913) | Eigenmann (1922) (Regan, 1913) “58 On ie Andinoacara latifrons x |x Steindachner (1878) | Maldonado-Ocam- (Steindachner, 1878) po et al. (2013b) | Eigenmann (1922) Andinoacara sapayensis (Regan, 1903) Eigenmann (1922) Bujurquina mariae (Eisenmann 1922) me | x Eigenmann (1922) Bujurquina moriorum Kullander, 1986 Rallanider (1286) Bujurquina peregrinabunda Kullander, 1986 Kallander (1986) ioe papel Gates | Mojjica (1999) | ICN-MHN 1523, 1529 oe EDe Tena es SADN Nog | a5 x Dahl (1960c) | ICN-MHN 95, 95a Cichlasoma amazonarum Kullander, 1983 a pura Cichl ) Eigenmann (1922) | FMNH 58614 [ex a ee pee x | x x CM 7639], 58615 [ex CM 7639] | CAS aria 66956 [ex IU 14171] Add cumin Kullander (1983) | ANSP 127364, A GRAY aa ae a 127364, 127397 | FMNH 84008 | NRM Kullander, 1983 39282 Laetacara flavilabris (Cope, 1870) xX IMCN 6026 *Laetacara fulvipinnis Staeck & Schindler, 2007 AUT GIRO? Laetacara thayeri bs (Steindachner, 1875) iMeeachalaoMg>) Caquetaia kraussii : , (Steindachner, 1879) X_ |Steindachner (1878) | Eigenmann (1922) Caquetaia myersi (Schultz, 1944) Schultz (1944c) | USNM 120533, 120534 100 Carlos DoNascimiento et al. / ZooKeys 708: 25-138 (2017) . | Mag- ; Taxa DC |EN| ET | Amz| Ori Ge Pac| Car| References/Collections Heroina isonycterina x Kullander (1996) Kullander, 1996 ae Heros efasciatus Heckel, 1840 Kullander (1986) Heros severus Heckel, 1840 Correa (2003) | Cala (1977) ; Bogota-Gregory and Maldonado- ee Be le X | X Ocampo (2006a) | Maldonado-Ocampo et al. (2006a) | [AVH-P 2224 Hypselecara coryphaenoides x |x Arbeléez et al. (2004) | Galvis et al. (Heckel, 1840) (2007a) Hypselecara temporalis (Giinther, 1862) x Kullander (1986) Kronoheros umbrifer (Meek & Hildebrand, 1913) A nal Pics Nite Ret aay all Se X | X [Regan (1912a) | Eigenmann (1922) (Regan, 1912) Mesoheros ornatus (Regan, 1905) Mesonauta egregius Kullander & Silfvergrip, 1991 Mesonauta insignis (Heckel, 1840) Mesonauta mirificus Kullander & Silfvergrip, 1991 + |Fowler (1944) | Eigenmann (1922) Kullander and Silfvergrip (1991) | FMNH 92863 | ICNMNH 1686 [ex NRM 43043] | NRM 11302, 12280 Kullander (1986) | Kullander and Silfvergrip (1991) Kullander and Silfvergrip (1991) Pterophyllum altum Pellegrin, 1903 Pterophyllum scalare (Schultze, 1823) Symphysodon aequifasciatus Pellegrin, 1904 Cala (1977) Arbeldez et al. (2004) Correa (2003) Uaru amphiacanthoides Heckel, 1840 Uaru fernandezyepezi Stawikowski, 1989 Beloniformes 4 spp (1 deleted from 2008) Correa (2003) Galvis et al. (2007a) Belonidae Belonion dibranchodon Collette, 1966 Potamorrhaphis guianensis (Jardine, 1843) Potamorrhaphis petersi Collette, 1974 ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ Correa (2003) | Maldonado-Ocampo (2001) Galvis et al. (2007b); Cala (1977) Galvis et al. (2007a) Pseudotylosurus microps (Giinther, 1866) Cyprinodontiformes 42 spp (7 added and 3 deleted from 2008) ~ ey eee ~ Mojica et al. (2005) | Maldonado- Ocampo (2001) Checklist of the freshwater fishes of Colombia: a Darwin Core alternative... 101 . | Mag- ; Taxa DC|EN| ET | Amz| Ori Ge Pac) Car | References/Collections Cynolebiidae 24 spp (5 added and 2 deleted from 2008) Anablepsoides atratus (Garman, 1895) xX NRM 14686 Anablepsoides elongatus (Fels & de Rham, 1981) x Correa (2003) Anablepsoides ophiomimus abe 193 lh I eae Anablepsoides ornatus . (Garman, 1895) xX Mojica et al. (2005) Anablepsoides rubrolineatus e (Fels & de Rham, 1981) x Mojica et al. (2005) Anablepsoides taeniatus Anablepsoides tessellatus (Huber, 1992) | x Huber (1992) | ANSP 139468, 168979 Austrofundulus guajira Hrbek, Taphorn & x X |Hrbek et al. (2005) Thomerson, 2005 Austrofundulus myersi Dahl, x |x x Dahl (1958) | SU 49513, 49513, 68324 | 1958 DZVU [ex SU 49513] Cynodonichthys boehlkei x |x Huber and Fels (1985) | ANSP 139467, (Huber & Fels, 1985) 139467 Cynodonichthys elegans x |x Steindachner (1880) | NMW 605441, (Steindachner, 1880) 605442-3, 18893-6 Cynodonichthys leucurus x | X x Fowler (1944) | ANSP 71436, 71437-50 (Fowler, 1944) Henn (1916) | CAS 44227 [ex CM 5814a-m | IU 13603], 52234 [ex IU phe Hee tena 6| X |X Xx 13606], 52235 [ex IU 13605], 78377 eae a [ex IU 13606] | FMNH 56997 [ex CM 5813], 56998-99, 57000-02 Cynodonichthys pacificus x |x x |x Huber (1992) | Maldonado-Ocampo et (Huber, 1992) al. (2013b) oe BP PEBEATIOIEE | oe 1 x Huber (1992) | NRM 16548, 14687 Laimosemion corpulentus Thomerson and Taphorn (1993) | SU (Thomerson & Taphorn, x | x x Inot CAS] 69692, 53693 1993) Laimosemion leticia ae Valdesalici, 2016 X |X x Valdesalici (2016) Ge TeV NB Set x X |Regan (1912d) | Mojica et al. (2006b) Rachovia hummelincki de Beaufort, 1940 (nee Taphorn and Thomerson (1978) Rachovia maculipinnis (Radda, 1964) x Taphorn and Thomerson (1978) Rivulus azurescens Vermeulen, 2013 x |X X Vermeulen (2013) 102 Carlos DoNascimiento et al. / ZooKeys 708: 25-138 (2017) . | Mag- ; Taxa DC|EN|ET | Amz| Ori ene Pac| Car | References/Collections Rivulus pivijay Vermeulen, x |x x Vermeulen (2013) 2013 Rivulus ribesrubrum Vermeulen 2013 4 X Vermeulen (2013) Rivulus xi Vermeulen, 2013 X X Vermeulen (2013) Cyprinodontidae 1 sp ** Vssolebias martae ‘ (Steindachner, 1876) X | xX eee Steindachner (1876) Poeciliidae 17 spp (2 added and 1 deleted from 2008) Fluviphylax obscurus Costa, x IAVH-P 2171, 2219 1996 Fluviphylax pygmaeus (Myers & Carvalho, 1955) Lucinda (2003) | NRM 26248 Gambusia lemaitrei Fowler, 1950 xX |X Fowler (1950) | ANSP 71938, 71939-40 Henn (1916) | FMNH 57007 [ex CM . 5823], 57008 [ex CM 5824a-g] | AMNH Ne eee ent oe i X |22705sw [ex FMNH 57008], 36481 [ex IU 13612] | CAS 22768 [ex IU 13612] | UMMZ 65232 [ex IU 13612] Poeciliopsis turrubarensis (Meck, 1912) LL |e) freeman ty Priapichthys caliensis x |x Henn (1916) | FMNH 57721 [ex CM (Eigenmann & Henn, 1916) 6700a], FMNH [ex CM 6700b] De eR ak Henn (1916) | CAS 22547 [ex IU OH tt f a é ew eal lake 13618], 22560 [ex IU 13619] | FMNH ie 57009 | UMMZ 65223 [ex IU 13619] *Priapichthys darienensis (Meek & Hildebrand, 1913) alban ae Priapichthys nigroventralis (Figenmann & Henn, 1912) | 2X X_ |Eigenmann (1912) | Henn (1916) Tey, Radda (1987) | FMNH 56045 [ex CM seed Loe ; ON, slice 4835], 54046 | BMNH 1924.3.3.85-86 ae a = | CAS 22765 [ex IU 12689] | USNM 79205 Pseudopoecilia fria Maldonado-Ocampo et al. (2013b) | (Eigenmann & Henn, 1914) ICN-MHN 2335 Poecilia caucana Steindachner (1880) | Mojica et al. (Steindachner, 1880) (2006b) *Poecilia gillii (Kner, 1863) X | Poeser (2003a) **Poecilia koperi Poeser, 2003 X | Poeser (2003b) Poecilia mechthildae Meyer, x |x x Meyer et al. (2002) | MTDF 26473- Etzel & Bork, 2002 26476 | SMF 28889 Poecilia orri Fowler, 1943 X_ |Poeser (2003a) Poecilia sphenops Valen cicninese THN ay) |) le Agudelo-Zamora et al. (2010) Checklist of the freshwater fishes of Colombia: a Darwin Core alternative... 103 Taxa DC|EN| ET | Amz) Ori eh Pac] Car| References/Collections Eupercaria incertae sedis 7 spp Sciaenidae Pachypops fourcroi (Lacepéde, 1802) Pachypops trifilis (Miiller & Troschel, 1849) Casatti (2002) | [AVH-P Bogota-Gregory and Maldonado-Ocam- po (2005) Pachyurus gabrielensis Casatti, 2001 Pachyurus junki Soares & x Maldonado-Ocampo et al. (2008) Bogota-Gregory and Maldonado-Ocam- Casatti, 2000 po (2006a) Pachyurus schomburgkii Giinther, 1860 Plagioscion magdalenae (Steindachner, 1878) Mojica et al. (2005) | Galvis et al. (2007a) Steindachner (1878) | Mojica et al. (2006b) Plagioscion squamosissimus al Mojica et al. (2005) | Cala (1977) | (Heckel, 1840) Casatti (2005) Tetraodontiformes | Tetraodontidae eee Mojica et al. (2005) Ceratodontiformes Lepidosirenidae Lepidosiren paradoxa Mojica et al. (2005) | Bogota-Gregory and Fitzinger, 1837 Maldonado-Ocampo (2006b) Total of additions by hydrographic system Total of deletions by hidrographic system | 44 | 32 | 37 | 24| 19 Total number of modifications by hidrographic system 240) alee Final remarks Confusions and misinterpretations related to the freshwater fishes distributed in Co- lombian territory and erroneous name assignations were included in previous checklists at local, regional or national levels, as pointed out in previous sections. The exhaus- tive verification process of the current checklist of the freshwater fishes of Colombia is a step forward linked to the ichthyology collections database depuration process in progress. These efforts will bring new opportunities to insert this information into new analysis dealing with new conservation opportunities or approaches to counteract the drivers of biodiversity decline of the Colombian freshwater fish fauna. New protected areas or portfolios have been proposed for freshwaters ecosystems in the Orinoco, Magdalena-Cauca and Amazon River basins in the recent years (Lasso et al. 2011b; Téllez et al. 2011; Portocarrero-Aya and Cowx 2016). However, some species level information taken into account to designate those areas were wrong; criteria like species richness or endemic species have to be re-evaluated based on the information 104 Carlos DoNascimiento et al. / ZooKeys 708: 25-138 (2017) provided in the current checklist. Expert criteria, although important on those analysis and proposals, also have to be reevaluated in light of rigorous data depuration at taxo- nomic and geographic levels (e.g. Herrera 2015). Applying a decision theory approach to guide and improve the process in the field of conservation planning is important to deal with an incomplete geographic coverage of taxonomic groups (Ornellas et al. 2011), as is the case of the Colombian freshwater fishes. As it was mentioned in the introduction, new scenarios are in discussion in refer- ence to new approaches dealing with the conservation of the freshwater ecosystems and their biota worldwide (Ormerod 2014). Colombia as a country with one of the richest freshwater fish faunas, faces enormous challenges due to the past and current human impacts that have been affecting the river systems running through the Colom- bian landscape. Most remarkable is that an important number of species have small distribution ranges in many trans-Andean rivers and piedmont areas of the Orinoco and Amazon regions. Finally, the data depuration process, conducted mainly at the species level, reinforces the importance of biological collections as repositories of the biodiversity patrimony and allows specimen identifications to be rechecked and verified. More efforts are needed by the institutions in charge of the Colombian ichthyological collections to guarantee the adequate conservation of their specimens. A continuous interinstitutional effort and collection support is an important task to be facilitated by ACICTIOS and the Colombian Biodiversity Information System (SiB Colombia). Those interactions have an enormous impact in terms of the need to maintain a continuous and rigorous updating list process. Moving forward (in addition to field work to fill gaps in poorly sampled basins), we identify as the main tasks remaining: 1) to continue the process of verification of fish species distributions at the sub-basin level or smaller biogeographic areas; and 2) to conduct revisionary taxonomic studies of specific species-rich fish groups found in Colombia. Ideally, these studies have to be developed in the context of an integrative taxonomic approach. Acknowledgements We thank the Asociacié6n Colombiana de Ictidlogos (ACICTIOS), Instituto de Inves- tigacién de Recursos Bioldgicos Alexander von Humboldt, The Nature Conservancy, Conservation International Colombia, WWF Colombia, Universidad de Antioquia, and Pontificia Universidad Javeriana for the support and funds provided throughout the project: “Catalogue of the Freshwater Fishes of Colombia”. For comments and grammar corrections on the manuscript we thank D. C. Taphorn. We gratefully ac- knowledge F.C. T. Lima and Sven O. Kullander who made a thorough revision of the Characidae and Bryconidae (FCTL) and Cyprinodontiformes and Cichliformes sections (SOK) of the checklist. We are indebted to A. Acosta (CIACOL), J. G. Al- bornoz-Garzén (CZUT-IC), and S. Prada-Pedreros and A. Urbano-Bonilla (MPUJ) for facilitating information of records in their respective collections. FAVN thanks Checklist of the freshwater fishes of Colombia: a Darwin Core alternative... 105 the Oficina de Investigaciones y Desarrollo Cientifico de la Universidad del Tolima, CORTOLIMA, and Grupo de Investigacién en Zoologia de la Universidad del Toli- ma for funding, logistic, and technical support. References Acero PA, Betancur-R R (2006) Real identity of the northern Colombian endemic sea catfish Galeichthys bonillai Miles, 1945 (Siluriformes: Ariidae). Cybium 30: 215-219. Acosta-Santos A, Agudelo E, Duque SR (2016) Diversidad de rayas de agua dulce en el rio Amazonas colombiano: resultados preliminares (Capitulo 14). 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Ministerio de Ambiente y Desarrollo Sostenible, WWF Colombia, Bogota, 486 pp. Checklist of the freshwater fishes of Colombia: a Darwin Core alternative... 137 Zarske A (2011) Beitrage zur Kenntnis der Vertreter der Gattungen Pyrrhulina Valenciennes, 1846 und Copella Myers, 1956 des nordéstlichen Siidamerika (Teleostei: Characiformes: Lebiasinidae). Vertebrate Zoology 61: 13-45. Zarske A (2015) Hemigrammus rubrostriatus spec. nov. - ein neuer Salmler aus Kolumbien und Revalidierung von Hemigrammus falsus Meinken, 1958 (Teleostei: Ostariophysi: Characi- dae). Vertebrate Zoology 65: 3-14. Zarske A, Géry J (2002) Der Blaurote Kolumbien-Salmler. Hyphessobrycon columbianus n. sp. - ein neuer Salmler (Teleostei, Characiformes, Characidae) aus dem kolumbianischen Darien. Das Aquarium: Monatsmagazin fiir Vivaristik, Bornheim 36: 22-30. Supplementary material | Checklist of the native freshwater fishes of Colombia Authors: Carlos DoNascimiento, Edgar Esteban Herrera-Collazos, Guido A. Herrera- R, Armando Ortega-Lara, Francisco A. Villa-Navarro, José Saulo Usma Oviedo, Javier A. Maldonado-Ocampo Data type: Microsoft Excel Worksheet (.xlsx) Copyright notice: This dataset is made available under the Open Database License (http://opendatacommons.org/licenses/odbl/1.0/). The Open Database License (ODDbL) is a license agreement intended to allow users to freely share, modify, and use this Dataset while maintaining this same freedom for others, provided that the original source and author(s) are credited. Link: https://doi.org/10.3897/zookeys.708.13897.suppl1 Supplementary material 2 New species described from Colombia after Maldonado-Ocampo et al. (2008) Authors: Carlos DoNascimiento, Edgar Esteban Herrera-Collazos, Guido A. Herrera- R, Armando Ortega-Lara, Francisco A. Villa-Navarro, José Saulo Usma Oviedo, Javier A. Maldonado-Ocampo Data type: Microsoft Word Document (.docx) Copyright notice: This dataset is made available under the Open Database License (http://opendatacommons.org/licenses/odbl/1.0/). The Open Database License (ODDbL) is a license agreement intended to allow users to freely share, modify, and use this Dataset while maintaining this same freedom for others, provided that the original source and author(s) are credited. Link: https://doi.org/10.3897/zookeys.708.13897.suppl2 138 Carlos DoNascimiento et al. / ZooKeys 708: 25-138 (2017) Supplementary material 3 New records of species for Colombia after Maldonado-Ocampo et al. (2008) Authors: Carlos DoNascimiento, Edgar Esteban Herrera-Collazos, Guido A. Herrera- R, Armando Ortega-Lara, Francisco A. Villa-Navarro, José Saulo Usma Oviedo, Javier A. Maldonado-Ocampo Data type: Microsoft Word Document (.docx) Copyright notice: This dataset is made available under the Open Database License (http://opendatacommons.org/licenses/odbl/1.0/). The Open Database License (ODDbL) is a license agreement intended to allow users to freely share, modify, and use this Dataset while maintaining this same freedom for others, provided that the original source and author(s) are credited. Link: https://doi.org/10.3897/zookeys.708.13897.suppl3 Supplementary material 4 Species listed in Maldonado-Ocampo et al. (2008) with geographic distribution corrected. Authors: Carlos DoNascimiento, Edgar Esteban Herrera-Collazos, Guido A. Herrera- R, Armando Ortega-Lara, Francisco A. Villa-Navarro, José Saulo Usma Oviedo, Javier A. Maldonado-Ocampo Data type: Microsoft Word Document (.docx) Copyright notice: This dataset is made available under the Open Database License (http://opendatacommons.org/licenses/odbl/1.0/). The Open Database License (ODbL) is a license agreement intended to allow users to freely share, modify, and use this Dataset while maintaining this same freedom for others, provided that the original source and author(s) are credited. Link: https://doi.org/10.3897/zookeys.708.13897.suppl4 Supplementary material 5 Species deleted from the Colombian continental ichthyofauna Authors: Carlos DoNascimiento, Edgar Esteban Herrera-Collazos, Guido A. Herrera- R, Armando Ortega-Lara, Francisco A. Villa-Navarro, José Saulo Usma Oviedo, Javier A. Maldonado-Ocampo Data type: Microsoft Word Document (.docx) Copyright notice: This dataset is made available under the Open Database License (http://opendatacommons.org/licenses/odbl/1.0/). The Open Database License (ODDbL) is a license agreement intended to allow users to freely share, modify, and use this Dataset while maintaining this same freedom for others, provided that the original source and author(s) are credited. Link: https://doi.org/10.3897/zookeys.708.13897.suppl5